About the Authors

Yucong Huang

Roles Data curation, Investigation, Methodology, Writing – original draft

Affiliation Guangdong Provincial Key Laboratory of Aquatic Animal Disease Control and Healthy culture & Key Laboratory of Control for Diseases of Aquatic Economic Animals of Guangdong Higher Education Institutes, Fisheries College of Guangdong Ocean University, Southern Marine Science and Engineering Guangdong Laboratory (Zhanjiang), Zhanjiang, GuangDong, China

Heng Sun

Contributed equally to this work with: Heng Sun, Jun Dong, Jianrong Zhang, Haoyu Wang, Lindi Yang, Yanping Li

Roles Investigation, Validation

Affiliation Guangdong Provincial Key Laboratory of Aquatic Animal Disease Control and Healthy culture & Key Laboratory of Control for Diseases of Aquatic Economic Animals of Guangdong Higher Education Institutes, Fisheries College of Guangdong Ocean University, Southern Marine Science and Engineering Guangdong Laboratory (Zhanjiang), Zhanjiang, GuangDong, China

Jun Dong

Contributed equally to this work with: Heng Sun, Jun Dong, Jianrong Zhang, Haoyu Wang, Lindi Yang, Yanping Li

Roles Validation

Affiliation Guangdong Provincial Key Laboratory of Aquatic Animal Disease Control and Healthy culture & Key Laboratory of Control for Diseases of Aquatic Economic Animals of Guangdong Higher Education Institutes, Fisheries College of Guangdong Ocean University, Southern Marine Science and Engineering Guangdong Laboratory (Zhanjiang), Zhanjiang, GuangDong, China

Jianrong Zhang

Contributed equally to this work with: Heng Sun, Jun Dong, Jianrong Zhang, Haoyu Wang, Lindi Yang, Yanping Li

Roles Validation

Affiliation Guangdong Provincial Key Laboratory of Aquatic Animal Disease Control and Healthy culture & Key Laboratory of Control for Diseases of Aquatic Economic Animals of Guangdong Higher Education Institutes, Fisheries College of Guangdong Ocean University, Southern Marine Science and Engineering Guangdong Laboratory (Zhanjiang), Zhanjiang, GuangDong, China

Haoyu Wang

Contributed equally to this work with: Heng Sun, Jun Dong, Jianrong Zhang, Haoyu Wang, Lindi Yang, Yanping Li

Roles Validation

Affiliation Guangdong Provincial Key Laboratory of Aquatic Animal Disease Control and Healthy culture & Key Laboratory of Control for Diseases of Aquatic Economic Animals of Guangdong Higher Education Institutes, Fisheries College of Guangdong Ocean University, Southern Marine Science and Engineering Guangdong Laboratory (Zhanjiang), Zhanjiang, GuangDong, China

Lindi Yang

Contributed equally to this work with: Heng Sun, Jun Dong, Jianrong Zhang, Haoyu Wang, Lindi Yang, Yanping Li

Roles Project administration

Affiliation Guangdong Provincial Key Laboratory of Aquatic Animal Disease Control and Healthy culture & Key Laboratory of Control for Diseases of Aquatic Economic Animals of Guangdong Higher Education Institutes, Fisheries College of Guangdong Ocean University, Southern Marine Science and Engineering Guangdong Laboratory (Zhanjiang), Zhanjiang, GuangDong, China

Yanping Li

Contributed equally to this work with: Heng Sun, Jun Dong, Jianrong Zhang, Haoyu Wang, Lindi Yang, Yanping Li

Roles Software

Affiliation Guangdong Provincial Key Laboratory of Aquatic Animal Disease Control and Healthy culture & Key Laboratory of Control for Diseases of Aquatic Economic Animals of Guangdong Higher Education Institutes, Fisheries College of Guangdong Ocean University, Southern Marine Science and Engineering Guangdong Laboratory (Zhanjiang), Zhanjiang, GuangDong, China

Yifan Wang

Roles Conceptualization, Writing – review & editing

beckvanm7@163.com

Affiliations Guangdong Provincial Key Laboratory of Aquatic Animal Disease Control and Healthy culture & Key Laboratory of Control for Diseases of Aquatic Economic Animals of Guangdong Higher Education Institutes, Fisheries College of Guangdong Ocean University, Southern Marine Science and Engineering Guangdong Laboratory (Zhanjiang), Zhanjiang, GuangDong, China, College of Veterinary Medicine, Southwest University, Chongqing, China

Competing Interests

The authors have declared that no competing interests exist.