About the Authors

Atsuko Ota

Roles Data curation, Formal analysis, Investigation, Methodology, Project administration, Validation, Visualization, Writing – original draft

Affiliation Aderans Company Limited, Shinjuku, Tokyo, Japan

Hiroaki Kitamura

Roles Data curation, Formal analysis, Investigation, Visualization, Writing – original draft

Affiliation Aderans Company Limited, Shinjuku, Tokyo, Japan

Keigo Sugimoto

Roles Data curation, Formal analysis, Investigation, Visualization, Writing – original draft

Affiliation Aderans Company Limited, Shinjuku, Tokyo, Japan

Miho Ogawa

Roles Conceptualization, Project administration, Resources, Writing – original draft

Affiliations OrganTech Inc., Chuo-ku, Tokyo, Japan, Laboratory for Organ Regeneration, RIKEN Center for Biosystems Dynamics Research (BDR), Kobe, Hyogo, Japan

Naoshi Dohmae

Roles Investigation, Methodology, Resources, Validation

Affiliation Biomolecular Characterization Unit, RIKEN Center for Sustainable Resource Science, Wako, Saitama, Japan

Hiroki Okuno

Roles Investigation, Methodology, Resources, Validation

Affiliation RIKEN, Nishina Center for Accelerator-Based Science, Wako, Saitama, Japan

Kazuya Takahashi

Roles Investigation, Methodology, Resources, Validation

Affiliation RIKEN, Nishina Center for Accelerator-Based Science, Wako, Saitama, Japan

Kazutaka Ikeda

Roles Investigation, Methodology, Resources, Validation

Affiliation Department of Applied Genomics, Laboratory of Biomolecule Analysis, Kazusa DNA Research Institute, Kisarazu, Chiba, Japan

Tsutomu Tomita

Roles Investigation, Resources

Affiliation Biobank, National Cerebral and Cardiovascular Center, Suita, Osaka, Japan

Naoki Matsuoka

Roles Investigation, Resources

Affiliation Kobe City Medical Center General Hospital, Kobe, Hyogo, Japan

Kunitaka Matsuishi

Roles Investigation, Resources

Affiliation Kobe City Medical Center General Hospital, Kobe, Hyogo, Japan

Tetsuro Inokuma

Roles Investigation, Resources

Affiliation Kobe City Medical Center General Hospital, Kobe, Hyogo, Japan

Tohru Nagano

Roles Investigation, Resources

Affiliation Kobe City Medical Center General Hospital, Kobe, Hyogo, Japan

Makoto Takeo

Roles Conceptualization, Project administration, Supervision, Writing – review & editing

Affiliation Laboratory for Organ Regeneration, RIKEN Center for Biosystems Dynamics Research (BDR), Kobe, Hyogo, Japan

Takashi Tsuji

Roles Conceptualization, Funding acquisition, Methodology, Project administration, Supervision, Writing – review & editing

takashi.tsuji@riken.jp

Affiliations OrganTech Inc., Chuo-ku, Tokyo, Japan, Laboratory for Organ Regeneration, RIKEN Center for Biosystems Dynamics Research (BDR), Kobe, Hyogo, Japan

Competing Interests

I have read the journal’s policy and the authors of this manuscript have the following competing interests: This work was partially performed under the condition of an Invention Agreement among RIKEN, Aderans Co., Ltd., Adjuvant Cosme Japan Co., Ltd., Daikin Industries, Ltd., Nippon Life Insurance, Company, Shimadzu Corporation, Tosoh Corporation and Organ Technologies Inc. Takashi Tsuji was an adviser for Organ Technologies Inc. until 2020. This does not alter our adherence to PLOS ONE policies on sharing data and materials.