About the Authors

Qiumeng Zhang

Roles Writing – original draft

Affiliations Research Institute of Forestry, Chinese Academy of Forestry, Beijing, China, Key Laboratory of Tree Breeding and Cultivation, National Forestry and Grassland Administration, Beijing, China, Urban Forest Research Center, National Forestry and Grassland Administration, Beijing, China

Baoquan Jia

Roles Writing – review & editing

jiabaoquan2006@163.com

Affiliations Research Institute of Forestry, Chinese Academy of Forestry, Beijing, China, Key Laboratory of Tree Breeding and Cultivation, National Forestry and Grassland Administration, Beijing, China, Urban Forest Research Center, National Forestry and Grassland Administration, Beijing, China

Tong Li

Roles Methodology

Affiliations Research Institute of Forestry, Chinese Academy of Forestry, Beijing, China, Key Laboratory of Tree Breeding and Cultivation, National Forestry and Grassland Administration, Beijing, China, Urban Forest Research Center, National Forestry and Grassland Administration, Beijing, China

Wenrui Li

Roles Methodology

Affiliations Research Institute of Forestry, Chinese Academy of Forestry, Beijing, China, Key Laboratory of Tree Breeding and Cultivation, National Forestry and Grassland Administration, Beijing, China, Urban Forest Research Center, National Forestry and Grassland Administration, Beijing, China

Competing Interests

On behalf of all authors, disclose any competing interests that could be perceived to bias this work—acknowledging all financial support and any other relevant financial or non-financial competing interests. This does not alter our adherence to PLOS ONE policies on sharing data and materials.