About the Authors

Shwe Yee Win

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Visualization, Writing – original draft, Writing – review & editing

Affiliation Laboratory of Infectious Diseases, Department of Disease Control, Faculty of Veterinary Medicine, Hokkaido University, Kita-ku, Sapporo, Japan

Shiro Murata

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Validation, Visualization, Writing – original draft, Writing – review & editing

murata@vetmed.hokudai.ac.jp

Affiliations Laboratory of Infectious Diseases, Department of Disease Control, Faculty of Veterinary Medicine, Hokkaido University, Kita-ku, Sapporo, Japan, Department of Advanced Pharmaceutics, Faculty of Veterinary Medicine, Hokkaido University, Kita-ku, Sapporo, Japan

Sotaro Fujisawa

Roles Conceptualization, Formal analysis, Investigation, Writing – review & editing

Affiliation Laboratory of Infectious Diseases, Department of Disease Control, Faculty of Veterinary Medicine, Hokkaido University, Kita-ku, Sapporo, Japan

Hikari Seo

Roles Investigation, Writing – review & editing

Affiliation Laboratory of Infectious Diseases, Department of Disease Control, Faculty of Veterinary Medicine, Hokkaido University, Kita-ku, Sapporo, Japan

Jumpei Sato

Roles Investigation, Writing – review & editing

Affiliation Laboratory of Infectious Diseases, Department of Disease Control, Faculty of Veterinary Medicine, Hokkaido University, Kita-ku, Sapporo, Japan

Yoshinosuke Motai

Roles Investigation, Writing – review & editing

Affiliation Laboratory of Infectious Diseases, Department of Disease Control, Faculty of Veterinary Medicine, Hokkaido University, Kita-ku, Sapporo, Japan

Takumi Sato

Roles Investigation, Writing – review & editing

Affiliation Vaxxinova Japan K.K., Minato-ku, Tokyo, Japan

Eiji Oishi

Roles Resources, Supervision, Writing – review & editing

Affiliation Vaxxinova Japan K.K., Minato-ku, Tokyo, Japan

Akira Taneno

Roles Resources, Supervision, Writing – review & editing

Affiliation Vaxxinova Japan K.K., Minato-ku, Tokyo, Japan

Lat Lat Htun

Roles Investigation, Writing – review & editing

Affiliation Department of Pharmacology and Parasitology, University of Veterinary Science, Yezin, Nay Pyi Taw, Myanmar

Saw Bawm

Roles Investigation, Writing – review & editing

Affiliations Department of Pharmacology and Parasitology, University of Veterinary Science, Yezin, Nay Pyi Taw, Myanmar, Department of Livestock and Aquaculture Research, Ministry of Agriculture, Livestock and Irrigation, Nay Pyi Taw, Myanmar

Tomohiro Okagawa

Roles Investigation, Resources, Writing – review & editing

Affiliation Department of Advanced Pharmaceutics, Faculty of Veterinary Medicine, Hokkaido University, Kita-ku, Sapporo, Japan

Naoya Maekawa

Roles Investigation, Writing – review & editing

Affiliation Department of Advanced Pharmaceutics, Faculty of Veterinary Medicine, Hokkaido University, Kita-ku, Sapporo, Japan

Satoru Konnai

Roles Project administration, Resources, Supervision, Writing – review & editing

Affiliations Laboratory of Infectious Diseases, Department of Disease Control, Faculty of Veterinary Medicine, Hokkaido University, Kita-ku, Sapporo, Japan, Department of Advanced Pharmaceutics, Faculty of Veterinary Medicine, Hokkaido University, Kita-ku, Sapporo, Japan

Kazuhiko Ohashi

Roles Funding acquisition, Project administration, Resources, Supervision, Writing – review & editing

Affiliations Laboratory of Infectious Diseases, Department of Disease Control, Faculty of Veterinary Medicine, Hokkaido University, Kita-ku, Sapporo, Japan, Department of Advanced Pharmaceutics, Faculty of Veterinary Medicine, Hokkaido University, Kita-ku, Sapporo, Japan, International Affairs Office, Faculty of Veterinary Medicine, Hokkaido University, Kita-ku, Sapporo, Japan

Competing Interests

SM, SK, and KO are authors of a patent covering the materials and techniques described in this manuscript (EU patent EP15800403.6; Japanese patent, 2016-523581). TS, EO, and AT are employed by Vaxxinova Japan K. K., Tokyo, Japan. All other authors have no competing interests to declare. This does not alter our adherence to PLOS ONE policies on sharing data and materials.