About the Authors

Zhenyu Qiu

Contributed equally to this work with: Zhenyu Qiu, Shouqing Guo

Roles Data curation, Investigation, Project administration, Writing – original draft

Affiliations Northwest Minzu University, Biomedical Research Center, Gansu Tech Innovation Center of Animal Cell, Lanzhou, China, Life Science and Engineering College of Northwest Minzu University, Lanzhou, China

Shouqing Guo

Contributed equally to this work with: Zhenyu Qiu, Shouqing Guo

Roles Data curation, Investigation, Writing – original draft

Affiliations Northwest Minzu University, Biomedical Research Center, Gansu Tech Innovation Center of Animal Cell, Lanzhou, China, Life Science and Engineering College of Northwest Minzu University, Lanzhou, China

Geng Liu

Roles Formal analysis

Affiliations Northwest Minzu University, Biomedical Research Center, Gansu Tech Innovation Center of Animal Cell, Lanzhou, China, Life Science and Engineering College of Northwest Minzu University, Lanzhou, China

Mengyuan Pei

Roles Formal analysis

Affiliations Northwest Minzu University, Biomedical Research Center, Gansu Tech Innovation Center of Animal Cell, Lanzhou, China, Life Science and Engineering College of Northwest Minzu University, Lanzhou, China

Yuejiao Liao

Roles Methodology

Affiliations Northwest Minzu University, Biomedical Research Center, Gansu Tech Innovation Center of Animal Cell, Lanzhou, China, Life Science and Engineering College of Northwest Minzu University, Lanzhou, China

Jiamin Wang

Roles Formal analysis

Affiliations Northwest Minzu University, Biomedical Research Center, Gansu Tech Innovation Center of Animal Cell, Lanzhou, China, Northwest Minzu University, Biomedical Research Center, Key Laboratory of Biotechnology & Bioengineering of State Ethnic Affairs Commission, Lanzhou, China

Jiayou Zhang

Roles Validation

Affiliations National Engineering Technology Research Center of Combined Vaccines, Wuhan, China, China National Biotec Group Company Limited, Beijing, China

Di Yang

Roles Validation

Affiliations Northwest Minzu University, Biomedical Research Center, Gansu Tech Innovation Center of Animal Cell, Lanzhou, China, Northwest Minzu University, Biomedical Research Center, Key Laboratory of Biotechnology & Bioengineering of State Ethnic Affairs Commission, Lanzhou, China

Zilin Qiao

Roles Data curation, Investigation, Project administration, Writing – original draft

Affiliations Northwest Minzu University, Biomedical Research Center, Gansu Tech Innovation Center of Animal Cell, Lanzhou, China, Northwest Minzu University, Biomedical Research Center, Key Laboratory of Biotechnology & Bioengineering of State Ethnic Affairs Commission, Lanzhou, China

Zhuo Li

Roles Methodology, Writing – review & editing

Affiliations Northwest Minzu University, Biomedical Research Center, Gansu Tech Innovation Center of Animal Cell, Lanzhou, China, Northwest Minzu University, Biomedical Research Center, Key Laboratory of Biotechnology & Bioengineering of State Ethnic Affairs Commission, Lanzhou, China

Zhongren Ma

Roles Project administration

Affiliations Northwest Minzu University, Biomedical Research Center, Gansu Tech Innovation Center of Animal Cell, Lanzhou, China, Northwest Minzu University, Biomedical Research Center, Key Laboratory of Biotechnology & Bioengineering of State Ethnic Affairs Commission, Lanzhou, China

Zhenbin Liu

Roles Methodology, Writing – review & editing

yangxiaoming@sinopharm.com (XY); liuzhenbin6@163.com (ZL)

Affiliations Northwest Minzu University, Biomedical Research Center, Gansu Tech Innovation Center of Animal Cell, Lanzhou, China, Northwest Minzu University, Biomedical Research Center, Key Laboratory of Biotechnology & Bioengineering of State Ethnic Affairs Commission, Lanzhou, China

Xiaoming Yang

Roles Conceptualization, Supervision, Writing – review & editing

yangxiaoming@sinopharm.com (XY); liuzhenbin6@163.com (ZL)

Affiliations National Engineering Technology Research Center of Combined Vaccines, Wuhan, China, China National Biotec Group Company Limited, Beijing, China

Competing Interests

The authors declare no conflict of interest.