About the Authors

Satohiro Nakao

Contributed equally to this work with: Satohiro Nakao, Kotono Ito

Roles Data curation, Investigation, Methodology, Validation, Writing – original draft

Affiliation Division of Reproductive Engineering, Center for Animal Resources and Development, Kumamoto University, Kumamoto, Japan

Kotono Ito

Contributed equally to this work with: Satohiro Nakao, Kotono Ito

Roles Investigation, Methodology, Validation

Affiliation Division of Reproductive Engineering, Center for Animal Resources and Development, Kumamoto University, Kumamoto, Japan

Chihiro Sugahara

Roles Investigation, Methodology

Affiliation Division of Reproductive Engineering, Center for Animal Resources and Development, Kumamoto University, Kumamoto, Japan

Hitomi Watanabe

Roles Investigation, Writing – review & editing

Affiliation Laboratory of Integrative Biological Science, Institute for Life and Medical Sciences, Kyoto University, Kyoto, Japan

Gen Kondoh

Roles Investigation, Writing – review & editing

Affiliation Laboratory of Integrative Biological Science, Institute for Life and Medical Sciences, Kyoto University, Kyoto, Japan

Naomi Nakagata

Roles Investigation, Writing – review & editing

Affiliation Division of Reproductive Biotechnology and Innovation, Center for Animal Resources and Development, Kumamoto University, Kumamoto, Japan

Toru Takeo

Roles Funding acquisition, Investigation, Methodology, Writing – review & editing

takeo@kumamoto-u.ac.jp

Affiliation Division of Reproductive Engineering, Center for Animal Resources and Development, Kumamoto University, Kumamoto, Japan

Competing Interests

The authors have declared that no competing interests exist.