About the Authors

Nicholas J. Santistevan

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Writing – original draft, Writing – review & editing

Affiliations Department of Integrative Biology, University of Wisconsin, Madison, Wisconsin, United States of America, Department of Neuroscience, University of Wisconsin, Madison, Wisconsin, United States of America, Genetics Ph.D. Training Program, University of Wisconsin, Madison, Wisconsin, United States of America

Jessica C. Nelson

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Writing – review & editing

Current address: Department of Cell and Developmental Biology, University of Colorado, Anschutz Medical Campus, Boulder, Colorado, United States of America

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania, Philadelphia, PA, United States of America

Elelbin A. Ortiz

Roles Conceptualization, Data curation, Formal analysis, Methodology, Writing – review & editing

Affiliations Department of Cell and Developmental Biology, University of Pennsylvania, Philadelphia, PA, United States of America, Neuroscience Graduate Program, University of Pennsylvania, Philadelphia, PA, United States of America

Andrew H. Miller

Roles Formal analysis, Methodology, Writing – review & editing

Affiliations Department of Integrative Biology, University of Wisconsin, Madison, Wisconsin, United States of America, Department of Cell and Developmental Biology, University of Pennsylvania, Philadelphia, PA, United States of America, Neuroscience Ph.D. Training Program, University of Wisconsin, Madison, Wisconsin, United States of America

Dima Kenj Halabi

Roles Investigation

Affiliations Department of Integrative Biology, University of Wisconsin, Madison, Wisconsin, United States of America, Department of Neuroscience, University of Wisconsin, Madison, Wisconsin, United States of America

Zoë A. Sippl

Roles Investigation

Affiliation Department of Integrative Biology, University of Wisconsin, Madison, Wisconsin, United States of America

Michael Granato

Roles Funding acquisition, Methodology, Supervision, Writing – review & editing

granatom@pennmedicine.upenn.edu (MG); ygrinblat@wisc.edu (YG)

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania, Philadelphia, PA, United States of America

Yevgenya Grinblat

Roles Conceptualization, Funding acquisition, Supervision, Writing – original draft, Writing – review & editing

granatom@pennmedicine.upenn.edu (MG); ygrinblat@wisc.edu (YG)

Affiliations Department of Integrative Biology, University of Wisconsin, Madison, Wisconsin, United States of America, Department of Neuroscience, University of Wisconsin, Madison, Wisconsin, United States of America, Genetics Ph.D. Training Program, University of Wisconsin, Madison, Wisconsin, United States of America

Competing Interests

The authors have declared that no competing interests exist.