About the Authors

Annette Klomp

Roles Conceptualization, Data curation, Formal analysis, Investigation, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliations Interdisciplinary Graduate Program in Neuroscience, University of Iowa, Iowa City, Iowa, United States of America, Iowa Neuroscience Institute, University of Iowa, Iowa City, Iowa, United States of America, Department of Molecular Physiology and Biophysics, University of Iowa, Iowa City, Iowa, United States of America

Ryotaro Omichi

Roles Formal analysis, Investigation

Current address: Department of Otolaryngology-Head and Neck Surgery, Okayama University Graduate School of Medicine, Dentistry and Pharmaceutical Sciences, Okayama, Japan

Affiliations Department of Otolaryngology-Head and Neck Surgery, University of Iowa, Iowa City, Iowa, United States of America, Iowa Institute of Human Genetics, University of Iowa, Iowa City, Iowa, United States of America

Yoichiro Iwasa

Roles Formal analysis, Investigation

Current address: Department of Otorhinolaryngology, Shinshu University School of Medicine, Matsumoto, Japan

Affiliations Department of Otolaryngology-Head and Neck Surgery, University of Iowa, Iowa City, Iowa, United States of America, Iowa Institute of Human Genetics, University of Iowa, Iowa City, Iowa, United States of America

Richard J. Smith

Roles Funding acquisition, Resources, Supervision

Affiliations Iowa Neuroscience Institute, University of Iowa, Iowa City, Iowa, United States of America, Department of Molecular Physiology and Biophysics, University of Iowa, Iowa City, Iowa, United States of America, Department of Otolaryngology-Head and Neck Surgery, University of Iowa, Iowa City, Iowa, United States of America, Iowa Institute of Human Genetics, University of Iowa, Iowa City, Iowa, United States of America

Yuriy M. Usachev

Roles Funding acquisition, Resources, Supervision

Affiliations Iowa Neuroscience Institute, University of Iowa, Iowa City, Iowa, United States of America, Department of Neuroscience and Pharmacology, University of Iowa, Iowa City, Iowa, United States of America

Andrew F. Russo

Roles Funding acquisition, Resources, Supervision

Affiliations Iowa Neuroscience Institute, University of Iowa, Iowa City, Iowa, United States of America, Department of Molecular Physiology and Biophysics, University of Iowa, Iowa City, Iowa, United States of America, Department of Neurology, University of Iowa, Iowa City, Iowa, United States of America

Nandakumar S. Narayanan

Roles Investigation, Project administration, Resources

Affiliations Iowa Neuroscience Institute, University of Iowa, Iowa City, Iowa, United States of America, Department of Neurology, University of Iowa, Iowa City, Iowa, United States of America

Amy Lee

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Project administration, Resources, Supervision, Writing – original draft, Writing – review & editing

amy.lee1@austin.utexas.edu

Current address: Department of Neuroscience, University of Texas-Austin, Austin, TX, United States of America

Affiliations Iowa Neuroscience Institute, University of Iowa, Iowa City, Iowa, United States of America, Department of Molecular Physiology and Biophysics, University of Iowa, Iowa City, Iowa, United States of America, Department of Otolaryngology-Head and Neck Surgery, University of Iowa, Iowa City, Iowa, United States of America, Department of Neurology, University of Iowa, Iowa City, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.