About the Authors

Alexander T. Duckworth

Roles Conceptualization, Investigation, Methodology, Writing – original draft, Writing – review & editing

Affiliation Department of Biomolecular Chemistry, University of Wisconsin-Madison, Madison, WI, United States of America

Tricia A. Windgassen

Roles Conceptualization, Investigation, Methodology, Writing – review & editing

Affiliations Department of Biomolecular Chemistry, University of Wisconsin-Madison, Madison, WI, United States of America, Codexis Inc, Redwood City, CA, United States of America

James L. Keck

Roles Conceptualization, Funding acquisition, Supervision, Writing – original draft, Writing – review & editing

jlkeck@wisc.edu

Affiliation Department of Biomolecular Chemistry, University of Wisconsin-Madison, Madison, WI, United States of America

Competing Interests

The authors have read the journal’s policy and have the following competing interests: Tricia A Windgassen became a paid employee of Codexis, Inc. after the study was completed. There are no patents, products in development or marketed products associated with this research to declare. This does not alter our adherence to PLOS ONE policies on sharing data and materials.