About the Authors

Lisa K. Lauderdale

Roles Conceptualization, Formal analysis, Methodology, Writing – original draft, Writing – review & editing

lisa.lauderdale@czs.org

Affiliation Conservation Science and Animal Welfare Research, Chicago Zoological Society–Brookfield Zoo, Brookfield, IL, United States of America

K. Alex Shorter

Roles Formal analysis, Writing – review & editing

Affiliation Department of Mechanical Engineering, University of Michigan, Ann Arbor, MI, United States of America

Ding Zhang

Roles Formal analysis, Writing – review & editing

Affiliation Department of Mechanical Engineering, University of Michigan, Ann Arbor, MI, United States of America

Joaquin Gabaldon

Roles Formal analysis, Writing – review & editing

Affiliation Robotics Institute, University of Michigan, Ann Arbor, MI, United States of America

Jill D. Mellen

Roles Conceptualization, Formal analysis, Funding acquisition, Methodology, Writing – review & editing

Affiliation Biology Department, Portland State University, Portland, OR, United States of America

Michael T. Walsh

Roles Conceptualization, Funding acquisition, Methodology, Writing – review & editing

Affiliation Department of Comparative, Diagnostic & Population Medicine, College of Veterinary Medicine, University of Florida, Gainesville, FL, United States of America

Douglas A. Granger

Roles Conceptualization, Funding acquisition, Methodology, Writing – review & editing

Affiliation Institute for Interdisciplinary Salivary Bioscience Research, University of California, Irvine, CA, United States of America

Lance J. Miller

Roles Conceptualization, Formal analysis, Funding acquisition, Methodology, Writing – original draft, Writing – review & editing

Affiliation Conservation Science and Animal Welfare Research, Chicago Zoological Society–Brookfield Zoo, Brookfield, IL, United States of America

Competing Interests

The authors have read the journal’s policy, and the authors of the study have the following competing interests to declare: LM and LL are affiliated with the Chicago Zoological Society – Brookfield Zoo (an AZA and AMMPA accredited zoo), and Chicago Zoological Society provides a salary for LM. Before the study, JM was previously affiliated with Disney’s Animal Kingdom (an AZA accredited zoo). The Seas® Epcot® Walt Disney World® Resort, Dolphin Island – Resorts World Sentosa, and Loro Parque and Loro Parque Fundación provided funding to the Chicago Zoological Society for this study. This does not alter our adherence to PLOS ONE policies on sharing data and materials. There are no patents, products in development or marketed products associated with this research to declare.