About the Authors

Sam Barker

Roles Formal analysis, Investigation, Writing – original draft

Current address: Midatech Pharma Plc, Cardiff, United Kingdom

Affiliation Department of Biosciences, College of Life and Environmental Sciences, University of Exeter, Exeter, United Kingdom

Sarah V. Harding

Roles Conceptualization, Supervision, Writing – original draft, Writing – review & editing

Affiliation Defence Science and Technology Laboratory, Porton Down, Salisbury, United Kingdom

David Gray

Roles Conceptualization, Supervision, Writing – original draft

Affiliation Drug Discovery Unit, University of Dundee, Dundee, Scotland

Mark I. Richards

Roles Investigation

Affiliation Defence Science and Technology Laboratory, Porton Down, Salisbury, United Kingdom

Helen S. Atkins

Roles Conceptualization, Supervision, Writing – original draft

Affiliations Department of Biosciences, College of Life and Environmental Sciences, University of Exeter, Exeter, United Kingdom, Defence Science and Technology Laboratory, Porton Down, Salisbury, United Kingdom

Nicholas J. Harmer

Roles Conceptualization, Formal analysis, Investigation, Methodology, Supervision, Writing – original draft, Writing – review & editing

N.J.Harmer@exeter.ac.uk

Affiliations Department of Biosciences, College of Life and Environmental Sciences, University of Exeter, Exeter, United Kingdom, Living Systems Institute, Exeter, United Kingdom

Competing Interests

The authors note that author SB was employed by Phoremost Ltd and Midatech Plc after completing his work on this manuscript, but before the final manuscript was completed. These companies provided his salary but did not have any influence on the text or data of the manuscript, or the decision to publish. This does not alter our adherence to PLOS ONE policies on sharing data and materials. The authors declare no other competing interests.