About the Authors

Mathew Hepp

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Writing – original draft

Affiliation Department of Biological Sciences, Centre for Wildlife Ecology, Simon Fraser University, Burnaby, BC, Canada

Eirikur Palsson

Roles Formal analysis, Methodology, Software, Validation, Writing – review & editing

Affiliation Department of Biological Sciences, Simon Fraser University, Burnaby, BC, Canada

Sarah K. Thomsen

Roles Formal analysis, Methodology, Visualization, Writing – original draft, Writing – review & editing

Current address: Department of Integrative Biology, Oregon State University, Corvallis, OR, United States of America

Affiliation Department of Biological Sciences, Centre for Wildlife Ecology, Simon Fraser University, Burnaby, BC, Canada

David J. Green

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Validation, Writing – review & editing

djgreen@sfu.ca

Affiliation Department of Biological Sciences, Centre for Wildlife Ecology, Simon Fraser University, Burnaby, BC, Canada

Competing Interests

Cooper Beauchesne and Associates Ltd provided field accommodation and a food allowance for a graduate student and field assistant that collected data used in this paper in each year from 2008-2017. This does not alter our adherence to PLOS ONE policies on sharing data and materials.