About the Authors

Masataka Michigami

Roles Conceptualization, Investigation, Methodology, Writing – original draft

Affiliation Department of Biological Science, Osaka Prefecture University, Sakai, Osaka, Japan

Kentaro Takahashi

Roles Investigation, Writing – review & editing

Affiliation Department of Biological Science, Osaka Prefecture University, Sakai, Osaka, Japan

Haruna Yamashita

Roles Investigation, Writing – review & editing

Affiliation Department of Biological Science, Osaka Prefecture University, Sakai, Osaka, Japan

Zhengmao Ye

Roles Investigation, Methodology, Writing – review & editing

Affiliation Interprotein Corporation, Osaka, Japan

Ikuhiko Nakase

Roles Investigation, Methodology, Writing – review & editing

Affiliation Department of Biological Science, Osaka Prefecture University, Sakai, Osaka, Japan

Ikuo Fujii

Roles Conceptualization, Funding acquisition, Project administration, Supervision, Writing – review & editing

fujii@b.s.osakafu-u.ac.jp

Affiliation Department of Biological Science, Osaka Prefecture University, Sakai, Osaka, Japan

Competing Interests

Co-author ZY is employed by Interprotein Corporation. This does not alter our adherence to PLOS ONE policies on sharing data and materials.