About the Authors

Caroline M. Best

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Validation, Writing – original draft, Writing – review & editing

cbest@harper-adams.ac.uk

Affiliation Department of Veterinary Health & Animal Sciences, Harper Adams University, Newport, Shropshire, United Kingdom

Alison Z. Pyatt

Roles Conceptualization, Methodology, Supervision, Writing – review & editing

Affiliation Equine Department, Hartpury University, Hartpury, Gloucestershire, United Kingdom

Janet Roden

Contributed equally to this work with: Janet Roden, Malgorzata Behnke, Kate Phillips

Roles Conceptualization, Supervision, Writing – review & editing

Affiliation Department of Veterinary Health & Animal Sciences, Harper Adams University, Newport, Shropshire, United Kingdom

Malgorzata Behnke

Contributed equally to this work with: Janet Roden, Malgorzata Behnke, Kate Phillips

Roles Conceptualization, Supervision, Writing – review & editing

Affiliation Department of Veterinary Health & Animal Sciences, Harper Adams University, Newport, Shropshire, United Kingdom

Kate Phillips

Contributed equally to this work with: Janet Roden, Malgorzata Behnke, Kate Phillips

Roles Conceptualization, Supervision, Writing – review & editing

Affiliation Department of Veterinary Health & Animal Sciences, Harper Adams University, Newport, Shropshire, United Kingdom

Competing Interests

I have read the journal’s policy and the authors of this manuscript have the following competing interests: CMB has received funding from a commercial source, MSD Animal Health. However, this does not alter the authors’ adherence to PLOS ONE policies on sharing data and materials. There are no patents, products in development or marketed products associated with this research to declare.