About the Authors

Joel Crespo

Roles Formal analysis, Investigation, Methodology, Writing – original draft, Writing – review & editing

Affiliation Immunology Research, Lilly Research Laboratories, Eli Lilly and Company, Indianapolis, IN, United States of America

Yi Ting Koh

Roles Formal analysis, Investigation, Methodology, Writing – review & editing

Affiliation Immunology Research, Lilly Research Laboratories, Eli Lilly and Company, Indianapolis, IN, United States of America

Ningjie Hu

Roles Formal analysis, Writing – review & editing

Affiliation Immunology Research, Lilly Research Laboratories, Eli Lilly and Company, Indianapolis, IN, United States of America

Paul A. Moore

Roles Project administration, Writing – review & editing

Affiliation MacroGenics, Rockville, MD, United States of America

Ezio Bonvini

Roles Investigation, Methodology, Writing – review & editing

Affiliation MacroGenics, Rockville, MD, United States of America

Andrew L. Glasebrook

Roles Investigation, Methodology, Writing – review & editing

Affiliation Immunology Research, Lilly Research Laboratories, Eli Lilly and Company, Indianapolis, IN, United States of America

Andrea P. Martin

Contributed equally to this work with: Andrea P. Martin, Robert J. Benschop

Roles Conceptualization, Formal analysis, Investigation, Methodology, Project administration, Supervision, Writing – original draft, Writing – review & editing

Current address: Immunology and Respiratory Disease Research, Boehringer Ingelheim Pharmaceuticals, Inc., Ridgefield, CT, United States of America

Affiliation Immunology Research, Lilly Research Laboratories, Eli Lilly and Company, Indianapolis, IN, United States of America

Robert J. Benschop

Contributed equally to this work with: Andrea P. Martin, Robert J. Benschop

Roles Conceptualization, Investigation, Methodology, Project administration, Supervision, Writing – original draft, Writing – review & editing

benschop_robert@lilly.com

Affiliation Immunology Research, Lilly Research Laboratories, Eli Lilly and Company, Indianapolis, IN, United States of America

Competing Interests

The authors have read the journal’s policy and the authors of this manuscript have the following competing interests: the following authors were employed by Eli Lilly and Company at the time of conducting the research and may own stock: JC, YTK, NH, ALG, APM, and RJB. The following authors were employed by Macrogenics at the time of conducting the research and may own stock: PAM and EB. APM is an employee of Boehringer Ingelheim Pharmaceuticals Inc. However, this affiliation was not held at the time the study was conducted. This does not alter our adherence to PLOS ONE policies on sharing data and materials. There are no patents, products in development or marketed products associated with this research to declare.