About the Authors

Mark Rawling

Contributed equally to this work with: Mark Rawling, Andrew Foey, Daniel Merrifield

Roles Data curation, Formal analysis, Investigation, Methodology, Visualization, Writing – original draft

mark.rawling@plymouth.ac.uk

Affiliation School of Biological and Marine Sciences, University of Plymouth, Plymouth, United Kingdom

Eric Leclercq

Roles Writing – review & editing

‡ These authors also contributed equally to this work

Affiliation Lallemand SAS, Blagnac, France

Andrew Foey

Contributed equally to this work with: Mark Rawling, Andrew Foey, Daniel Merrifield

Roles Conceptualization, Writing – review & editing

Affiliation School of Biomedical Sciences, University of Plymouth, Plymouth, United Kingdom

Mathieu Castex

Roles Funding acquisition, Project administration, Resources, Writing – review & editing

‡ These authors also contributed equally to this work

Affiliation Lallemand SAS, Blagnac, France

Daniel Merrifield

Contributed equally to this work with: Mark Rawling, Andrew Foey, Daniel Merrifield

Roles Supervision, Writing – review & editing

Affiliation School of Biological and Marine Sciences, University of Plymouth, Plymouth, United Kingdom

Competing Interests

The study was supported financially by Lallemand SAS. The commercial affiliation with this study does not alter our adherence to PLOS ONE policies on sharing data and materials.