About the Authors

Hayao Kobayashi

Roles Data curation, Formal analysis, Investigation, Methodology, Visualization, Writing – original draft

physeter0809@gmail.com

Affiliation Graduate School of Fisheries and Environmental Science, Nagasaki University, Nagasaki, Japan

Hal Whitehead

Roles Supervision, Validation, Writing – review & editing

Affiliation Department of Biology, Dalhousie University, Halifax, NS, Canada

Masao Amano

Roles Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Validation, Writing – review & editing

Affiliation Graduate School of Fisheries and Environmental Science, Nagasaki University, Nagasaki, Japan

Competing Interests

The authors have declared that no competing interests exist.