About the Authors

Taneli Heikkinen

Contributed equally to this work with: Taneli Heikkinen, Timo Bragge

Roles Investigation, Methodology, Supervision, Writing – original draft, Writing – review & editing

Affiliation Charles River Discovery Services, Kuopio, Finland

Timo Bragge

Contributed equally to this work with: Taneli Heikkinen, Timo Bragge

Roles Conceptualization, Data curation, Formal analysis, Investigation, Software, Writing – original draft, Writing – review & editing

Affiliation Charles River Discovery Services, Kuopio, Finland

Niina Bhattarai

Roles Investigation

Affiliation School of Pharmacy, Faculty of Health Sciences, University of Eastern Finland, Kuopio, Finland

Teija Parkkari

Roles Project administration

Affiliation Charles River Discovery Services, Kuopio, Finland

Jukka Puoliväli

Roles Resources

Affiliation Charles River Discovery Services, Kuopio, Finland

Outi Kontkanen

Roles Project administration

Affiliation Charles River Discovery Services, Kuopio, Finland

Patrick Sweeney

Roles Resources

Affiliation Naason Science Inc., Chungcheongbuk-do, South Korea

Larry C. Park

Roles Conceptualization, Funding acquisition, Supervision, Writing – original draft

Affiliations Naason Science Inc., Chungcheongbuk-do, South Korea, CHDI Management/CHDI Foundation, Los Angeles, California, United States of America

Ignacio Munoz-Sanjuan

Roles Funding acquisition, Resources, Supervision, Writing – original draft, Writing – review & editing

ignacio.munoz@chdifoundation.org

Affiliation CHDI Management/CHDI Foundation, Los Angeles, California, United States of America

Competing Interests

CHDI Foundation is a nonprofit biomedical research organization exclusively dedicated to collaboratively developing therapeutics that substantially improve the lives of those affected by Huntington’s disease. CHDI Foundation conducts research in a number of different ways; for the purposes of this manuscript, all research was conceptualized, planned, and directed by all authors and conducted at the contract research organization Charles River Discovery Services Finland. This does not alter our adherence to PLOS ONE policies on sharing data and materials.