About the Authors

Masaki Kohno

Roles Formal analysis, Investigation, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliations Department of Biotechnology, The University of Tokyo, Tokyo, Japan, R&D Division, HAYASHIBARA CO., LTD., Okayama, Japan

Takatoshi Arakawa

Roles Data curation, Formal analysis, Investigation, Project administration, Resources, Writing – review & editing

Affiliations Department of Biotechnology, The University of Tokyo, Tokyo, Japan, Collaborative Research Institute for Innovative Microbiology, The University of Tokyo, Tokyo, Japan

Naoki Sunagawa

Roles Formal analysis, Investigation, Resources, Validation, Writing – review & editing

Affiliation Department of Biomaterial Sciences, The University of Tokyo, Tokyo, Japan

Tetsuya Mori

Roles Methodology, Resources, Writing – review & editing

Affiliation R&D Division, HAYASHIBARA CO., LTD., Okayama, Japan

Kiyohiko Igarashi

Roles Funding acquisition, Supervision, Validation, Writing – review & editing

Affiliations Department of Biomaterial Sciences, The University of Tokyo, Tokyo, Japan, VTT Technical Research Centre of Finland Ltd., Espoo, Finland

Tomoyuki Nishimoto

Roles Conceptualization, Project administration, Supervision, Writing – review & editing

Affiliation R&D Division, HAYASHIBARA CO., LTD., Okayama, Japan

Shinya Fushinobu

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Project administration, Supervision, Validation, Writing – original draft, Writing – review & editing

asfushi@mail.ecc.u-tokyo.ac.jp

Affiliations Department of Biotechnology, The University of Tokyo, Tokyo, Japan, Collaborative Research Institute for Innovative Microbiology, The University of Tokyo, Tokyo, Japan

Competing Interests

The authors of this manuscript have the following competing interests: Three authors (MK, TM,TN) are paid employees of Hayashibara Co, Ltd. This commercial affiliation does not alter our adherence to PLOS ONE policies on sharing data and materials. There are no patents, products in development or marketed products associated with this research to declare.