About the Authors

Thomas A. Jefferson

Contributed equally to this work with: Thomas A. Jefferson, Mari A. Smultea

Roles Formal analysis, Investigation, Methodology, Writing – original draft

sclymene@aol.com

Affiliation Clymene Enterprises, Lakeside, CA, United States of America

Mari A. Smultea

Contributed equally to this work with: Thomas A. Jefferson, Mari A. Smultea

Roles Funding acquisition, Project administration, Writing – original draft

Affiliation Smultea Environmental Sciences, Preston, WA, United States of America

Eric J. Ward

Roles Formal analysis, Methodology, Writing – review & editing

‡ These authors also contributed equally to this work.

Affiliation Conservation Biology Division, Northwest Fisheries Science Center, National Marine Fisheries Service, National Oceanographic and Atmospheric Administration, Seattle, WA, United States of America

Barry Berejikian

Roles Investigation, Writing – review & editing

‡ These authors also contributed equally to this work.

Affiliation Environmental and Fisheries Sciences Division, Northwest Fisheries Science Center, National Marine Fisheries Service, National Oceanographic and Atmospheric Administration, Port Orchard, WA, United States of America

Competing Interests

Two of the authors are associated with scientific consulting companies, Clymene Enterprises LLC [TAJ] and Smultea Environmental Sciences LLC [MS]. This does not alter our adherence to PLOS ONE policies on sharing data and materials.