About the Authors

Joel H. Nitta

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Visualization, Writing – original draft, Writing – review & editing

joelnitta@gmail.com

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Atsushi Ebihara

Roles Funding acquisition, Resources, Writing – review & editing

Affiliation Department of Botany, National Museum of Nature and Science, Tsukuba, Ibaraki, Japan

Alan R. Smith

Roles Conceptualization, Formal analysis, Methodology, Writing – review & editing

Affiliation The University Herbarium, University of California at Berkeley, Berkeley, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.