About the Authors

Nobuyasu Hirai

Roles Data curation, Investigation, Project administration, Writing – original draft

Affiliations Department of Pathogen, Infection and Immunity, Nara Medical University, Kashihara, Nara, Japan, Center for Infectious Diseases, Nara Medical University, Kashihara, Nara, Japan, Department of Microbiology and Infectious Diseases, Nara Medical University, Kashihara, Nara, Japan

Kei Kasahara

Roles Supervision, Writing – review & editing

kassan@naramed-u.ac.jp

Affiliation Center for Infectious Diseases, Nara Medical University, Kashihara, Nara, Japan

Ryuichi Nakano

Roles Methodology, Supervision

Affiliation Department of Microbiology and Infectious Diseases, Nara Medical University, Kashihara, Nara, Japan

Yoshihiko Ogawa

Roles Project administration, Supervision

Affiliation Center for Infectious Diseases, Nara Medical University, Kashihara, Nara, Japan

Yuki Suzuki

Roles Data curation, Methodology

Affiliation Department of Microbiology and Infectious Diseases, Nara Medical University, Kashihara, Nara, Japan

Miho Ogawa

Roles Methodology

Affiliation BML Biomedical Laboratories R&D Center, Inc., Kawagoe, Saitama, Japan

Naokuni Hishiya

Roles Methodology

Affiliation Department of Microbiology and Infectious Diseases, Nara Medical University, Kashihara, Nara, Japan

Akiyo Nakano

Roles Supervision, Validation

Affiliation Department of Microbiology and Infectious Diseases, Nara Medical University, Kashihara, Nara, Japan

Sadahiro Ichimura

Roles Methodology

Affiliation BML Biomedical Laboratories R&D Center, Inc., Kawagoe, Saitama, Japan

Hisakazu Yano

Roles Conceptualization, Project administration, Supervision

Affiliation Department of Microbiology and Infectious Diseases, Nara Medical University, Kashihara, Nara, Japan

Masahide Yoshikawa

Roles Conceptualization, Supervision

Affiliation Department of Pathogen, Infection and Immunity, Nara Medical University, Kashihara, Nara, Japan

Competing Interests

The authors have declared that no competing interests exist. BML Biomedical Laboratories R&D Center, Inc., provided support in the form of salaries for the author MO and SI, but did not have any additional role in the study design, data collection and analysis, decision to poublish, or preparation of the manuscript. This does not alter our adherence to PLOS ONE policies on sharing data and materials.