About the Authors

Keiko Kitagawa

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Writing – original draft, Writing – review & editing

keiko.kitagawa@uni-tuebingen.de

Affiliations SFB 1070 ResourceCultures, University of Tübingen, Tübingen, Germany, Department of Early Prehistory and Quaternary Ecology, University of Tübingen, Tübingen, Germany

Nicholas J. Conard

Roles Conceptualization, Funding acquisition, Resources, Supervision, Writing – review & editing

Affiliations Department of Early Prehistory and Quaternary Ecology, University of Tübingen, Tübingen, Germany, Senckenberg Centre for Human Evolution and Paleoenvironment, University of Tübingen, Tübingen, Germany

Competing Interests

Authors have declared that no competing interests exist. Heidelberger Cement and Voith Corporation provided funding for this study. This does not alter our adherence to PLOS ONE policies on data sharing and materials."