About the Authors

Zhanyang Li

Roles Data curation, Funding acquisition, Investigation, Project administration, Supervision

Affiliation Institute of Cultural Heritage, Shandong University, Qingdao, Shandong Province, P.R. of China

Luc Doyon

Contributed equally to this work with: Luc Doyon, Francesco d’Errico

Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Methodology, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliations Institute of Cultural Heritage, Shandong University, Qingdao, Shandong Province, P.R. of China, Centre National de la Recherche Scientifique CNRS UMR5199 PACEA, Université de Bordeaux, Pessac, Nouvelle-Aquitaine, France

Hui Fang

Roles Funding acquisition, Project administration, Resources

francesco.derrico@u-bordeaux.fr (FdE); fangh@sdu.edu.cn (HF)

Affiliation Institute of Cultural Heritage, Shandong University, Qingdao, Shandong Province, P.R. of China

Ronan Ledevin

Roles Data curation, Formal analysis, Methodology, Software, Writing – original draft

‡ These authors also contributed equally to this work.

Affiliation Centre National de la Recherche Scientifique CNRS UMR5199 PACEA, Université de Bordeaux, Pessac, Nouvelle-Aquitaine, France

Alain Queffelec

Roles Formal analysis, Investigation, Software, Writing – review & editing

‡ These authors also contributed equally to this work.

Affiliation Centre National de la Recherche Scientifique CNRS UMR5199 PACEA, Université de Bordeaux, Pessac, Nouvelle-Aquitaine, France

Emeline Raguin

Roles Formal analysis, Investigation, Methodology, Resources, Software, Writing – original draft, Writing – review & editing

‡ These authors also contributed equally to this work.

Affiliation Department of Structural Biology, Weizmann Institute of Science, Rehovot, Israel

Francesco d’Errico

Contributed equally to this work with: Luc Doyon, Francesco d’Errico

Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Methodology, Visualization, Writing – original draft, Writing – review & editing

francesco.derrico@u-bordeaux.fr (FdE); fangh@sdu.edu.cn (HF)

Current address: CNRS UMR5199 –PACEA, Université de Bordeaux, Allée Geoffroy Saint Hilaire, Pessac, France

Affiliations Centre National de la Recherche Scientifique CNRS UMR5199 PACEA, Université de Bordeaux, Pessac, Nouvelle-Aquitaine, France, SFF Centre for Early Sapiens Behaviour (SapienCE), University of Bergen, Bergen, Norway

Competing Interests

The authors have declared that no competing interests exist.