About the Authors

Yunqiu Shang

Contributed equally to this work with: Yunqiu Shang, Ping Li

Roles Conceptualization, Data curation, Formal analysis, Investigation, Writing – original draft

Affiliation College of Agronomy, Shandong Agricultural University, State Key Laboratory of Crop Biology, Key Laboratory of Crop Ecophysiology and Farming System, Ministry of Agriculture, Taian, Shandong, People’s Republic of China

Xiang Lin

Roles Conceptualization, Data curation, Methodology

Affiliation College of Agronomy, Shandong Agricultural University, State Key Laboratory of Crop Biology, Key Laboratory of Crop Ecophysiology and Farming System, Ministry of Agriculture, Taian, Shandong, People’s Republic of China

Ping Li

Contributed equally to this work with: Yunqiu Shang, Ping Li

Roles Data curation, Formal analysis, Investigation

Affiliation College of Agronomy, Shandong Agricultural University, State Key Laboratory of Crop Biology, Key Laboratory of Crop Ecophysiology and Farming System, Ministry of Agriculture, Taian, Shandong, People’s Republic of China

Shubo Gu

Roles Methodology

Affiliation College of Agronomy, Shandong Agricultural University, State Key Laboratory of Crop Biology, Key Laboratory of Crop Ecophysiology and Farming System, Ministry of Agriculture, Taian, Shandong, People’s Republic of China

Keyi Lei

Roles Investigation

Affiliation College of Agronomy, Shandong Agricultural University, State Key Laboratory of Crop Biology, Key Laboratory of Crop Ecophysiology and Farming System, Ministry of Agriculture, Taian, Shandong, People’s Republic of China

Sen Wang

Roles Investigation

Affiliation College of Agronomy, Shandong Agricultural University, State Key Laboratory of Crop Biology, Key Laboratory of Crop Ecophysiology and Farming System, Ministry of Agriculture, Taian, Shandong, People’s Republic of China

Xinhui Hu

Roles Investigation

Affiliation College of Agronomy, Shandong Agricultural University, State Key Laboratory of Crop Biology, Key Laboratory of Crop Ecophysiology and Farming System, Ministry of Agriculture, Taian, Shandong, People’s Republic of China

Panpan Zhao

Roles Investigation

Affiliation College of Agronomy, Shandong Agricultural University, State Key Laboratory of Crop Biology, Key Laboratory of Crop Ecophysiology and Farming System, Ministry of Agriculture, Taian, Shandong, People’s Republic of China

Dong Wang

Roles Conceptualization, Formal analysis, Methodology, Project administration, Writing – review & editing

wangd@sdau.edu.cn

Affiliations College of Agronomy, Shandong Agricultural University, State Key Laboratory of Crop Biology, Key Laboratory of Crop Ecophysiology and Farming System, Ministry of Agriculture, Taian, Shandong, People’s Republic of China, Zibo HeFeng Seed Technology co., ltd., Linzi, Zibo, Shandong, People’s Republic of China

Competing Interests

The author DW is part-time employed by and receives salary from Zibo HeFeng Seed Technology. This does not alter our adherence to PLOS ONE policies on sharing data and materials. There are no patents, products in development or marketed products associated with this research to declare. The other authors declare no other competing interests exist.