About the Authors

Xiyang Zhang

Roles Data curation, Formal analysis, Writing – original draft, Writing – review & editing

Affiliations Zhuhai Key Laboratory of Marine Bioresources and Environment, Guangdong Provincial Key Laboratory of Marine Resources and Coastal Engineering, School of Marine Sciences, Sun Yat-Sen University, Zhuhai, China, Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai, China

Xi Ning

Roles Data curation

Affiliations Zhuhai Key Laboratory of Marine Bioresources and Environment, Guangdong Provincial Key Laboratory of Marine Resources and Coastal Engineering, School of Marine Sciences, Sun Yat-Sen University, Zhuhai, China, Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai, China

Xiaoxiao He

Roles Writing – review & editing

Affiliations Zhuhai Key Laboratory of Marine Bioresources and Environment, Guangdong Provincial Key Laboratory of Marine Resources and Coastal Engineering, School of Marine Sciences, Sun Yat-Sen University, Zhuhai, China, Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai, China

Xian Sun

Roles Writing – review & editing

Affiliations Zhuhai Key Laboratory of Marine Bioresources and Environment, Guangdong Provincial Key Laboratory of Marine Resources and Coastal Engineering, School of Marine Sciences, Sun Yat-Sen University, Zhuhai, China, Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai, China

Xinjian Yu

Roles Data curation

Affiliations Zhuhai Key Laboratory of Marine Bioresources and Environment, Guangdong Provincial Key Laboratory of Marine Resources and Coastal Engineering, School of Marine Sciences, Sun Yat-Sen University, Zhuhai, China, Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai, China

Yuanxiong Cheng

Roles Data curation

Affiliation The Third Affiliated Hospital of Southern Medical University, Guangzhou, China

Ri-Qing Yu

Roles Supervision, Writing – review & editing

exwyp@mail.sysu.edu.cn (YW); ryu@uttyler.edu (RQY)

Affiliation Department of Biology, Center for Environment, Biodiversity and Conservation, The University of Texas at Tyler, Tyler, Texas, United States of America

Yuping Wu

Roles Funding acquisition, Supervision, Writing – review & editing

exwyp@mail.sysu.edu.cn (YW); ryu@uttyler.edu (RQY)

Affiliations Zhuhai Key Laboratory of Marine Bioresources and Environment, Guangdong Provincial Key Laboratory of Marine Resources and Coastal Engineering, School of Marine Sciences, Sun Yat-Sen University, Zhuhai, China, Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai, China

Competing Interests

The authors have declared that no competing interests exist.