About the Authors

Rosemary J. Steinbach

Roles Data curation, Formal analysis, Investigation, Project administration, Visualization, Writing – original draft, Writing – review & editing

Affiliation Division of Reproductive & Developmental Sciences, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Nicole N. Haese

Roles Data curation, Investigation, Visualization, Writing – original draft, Writing – review & editing

Affiliations Vaccine & Gene Therapy Institute, Oregon Health & Science University, Beaverton, Oregon, United States of America, Division of Pathobiology & Immunology, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Jessica L. Smith

Roles Data curation, Investigation

Affiliation Vaccine & Gene Therapy Institute, Oregon Health & Science University, Beaverton, Oregon, United States of America

Lois M. A. Colgin

Roles Data curation, Formal analysis, Investigation, Visualization, Writing – original draft, Writing – review & editing

Affiliation Division of Comparative Medicine, Pathology Services Unit, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Rhonda P. MacAllister

Roles Methodology, Supervision

Affiliation Division of Comparative Medicine, Clinical Medicine Unit, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Justin M. Greene

Roles Data curation, Investigation

Affiliations Vaccine & Gene Therapy Institute, Oregon Health & Science University, Beaverton, Oregon, United States of America, Division of Pathobiology & Immunology, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Christopher J. Parkins

Roles Data curation, Investigation

Affiliation Vaccine & Gene Therapy Institute, Oregon Health & Science University, Beaverton, Oregon, United States of America

J. Beth Kempton

Roles Data curation, Investigation, Visualization, Writing – original draft, Writing – review & editing

Affiliation Department of Otolaryngology, Oregon Hearing Research Center, Oregon Health & Science University, Portland, Oregon, United States of America

Edward Porsov

Roles Data curation, Investigation, Resources, Software, Validation

Affiliation Department of Otolaryngology, Oregon Hearing Research Center, Oregon Health & Science University, Portland, Oregon, United States of America

Xiaojie Wang

Roles Data curation, Formal analysis, Investigation, Visualization, Writing – original draft, Writing – review & editing

Affiliation Advanced Imaging Research Center, Oregon Health & Science University, Portland, Oregon, United States of America

Lauren M. Renner

Roles Data curation, Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Neuroscience, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Trevor J. McGill

Roles Investigation, Visualization, Writing – original draft, Writing – review & editing

Affiliations Department of Neuroscience, Oregon National Primate Research Center, Beaverton, Oregon, United States of America, Department of Ophthalmology, Casey Eye Institute, Oregon Health & Science University, Portland, Oregon, United States of America

Brandy L. Dozier

Roles Investigation, Methodology, Supervision, Writing – review & editing

Affiliation Division of Comparative Medicine, Clinical Medicine Unit, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Craig N. Kreklywich

Roles Data curation, Investigation, Writing – review & editing

Affiliation Vaccine & Gene Therapy Institute, Oregon Health & Science University, Beaverton, Oregon, United States of America

Takeshi F. Andoh

Roles Investigation

Affiliation Vaccine & Gene Therapy Institute, Oregon Health & Science University, Beaverton, Oregon, United States of America

Marjorie R. Grafe

Roles Data curation, Formal analysis

Affiliation Department of Pathology, Oregon Health & Science University, Portland, Oregon, United States of America

Heidi L. Pecoraro

Roles Data curation, Formal analysis

Affiliation Veterinary Diagnostic Services Department, North Dakota State University, Fargo, North Dakota, United States of America

Travis Hodge

Roles Investigation, Resources

Affiliation Division of Comparative Medicine, Time Mated Breeding Services Unit, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Robert M. Friedman

Roles Formal analysis, Investigation, Methodology, Writing – review & editing

Affiliation Department of Neuroscience, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Lisa A. Houser

Roles Investigation

Affiliation Division of Comparative Medicine, Behavioral Services Unit, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Terry K. Morgan

Roles Formal analysis, Writing – review & editing

Affiliations Department of Pathology, Oregon Health & Science University, Portland, Oregon, United States of America, Department of Obstetrics & Gynecology, Oregon Health & Science University, Portland, Oregon, United States of America

Peter Stenzel

Roles Formal analysis, Writing – review & editing

Affiliation Department of Pathology, Oregon Health & Science University, Portland, Oregon, United States of America

Jonathan R. Lindner

Roles Formal analysis, Investigation

Affiliation Knight Cardiovascular Institute, Oregon Health & Science University, Portland, Oregon, United States of America

Robert L. Schelonka

Roles Methodology, Supervision, Writing – review & editing

Affiliation Division of Neonatology, Department of Pediatrics, Oregon Health & Science University, Portland, Oregon, United States of America

Jonah B. Sacha

Roles Methodology, Supervision, Writing – review & editing

Affiliations Vaccine & Gene Therapy Institute, Oregon Health & Science University, Beaverton, Oregon, United States of America, Division of Pathobiology & Immunology, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Victoria H. J. Roberts

Roles Methodology, Writing – review & editing

Affiliation Division of Reproductive & Developmental Sciences, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Martha Neuringer

Roles Methodology, Supervision

Affiliations Department of Neuroscience, Oregon National Primate Research Center, Beaverton, Oregon, United States of America, Department of Ophthalmology, Casey Eye Institute, Oregon Health & Science University, Portland, Oregon, United States of America

John V. Brigande

Roles Investigation, Methodology, Supervision, Writing – original draft, Writing – review & editing

Affiliation Department of Otolaryngology, Oregon Hearing Research Center, Oregon Health & Science University, Portland, Oregon, United States of America

Christopher D. Kroenke

Roles Methodology, Supervision

Affiliation Advanced Imaging Research Center, Oregon Health & Science University, Portland, Oregon, United States of America

Antonio E. Frias

Roles Investigation, Methodology, Supervision, Writing – review & editing

Affiliations Division of Reproductive & Developmental Sciences, Oregon National Primate Research Center, Beaverton, Oregon, United States of America, Department of Obstetrics & Gynecology, Oregon Health & Science University, Portland, Oregon, United States of America

Anne D. Lewis

Roles Data curation, Formal analysis, Investigation, Supervision, Visualization, Writing – original draft, Writing – review & editing

Affiliation Division of Comparative Medicine, Pathology Services Unit, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Meredith A. Kelleher

Roles Methodology, Supervision, Writing – original draft, Writing – review & editing

Affiliation Division of Reproductive & Developmental Sciences, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Alec J. Hirsch

Roles Conceptualization, Data curation, Investigation, Methodology, Supervision, Writing – original draft, Writing – review & editing

Affiliations Vaccine & Gene Therapy Institute, Oregon Health & Science University, Beaverton, Oregon, United States of America, Division of Pathobiology & Immunology, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Daniel Neal Streblow

Roles Conceptualization, Funding acquisition, Investigation, Methodology, Supervision, Writing – original draft, Writing – review & editing

streblow@ohsu.edu

Affiliations Vaccine & Gene Therapy Institute, Oregon Health & Science University, Beaverton, Oregon, United States of America, Division of Pathobiology & Immunology, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Competing Interests

The authors have declared that no competing interests exist.