About the Authors

Saeideh Esmaeili

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Visualization, Writing – original draft

saeideh.esmaeili@uwyo.edu

Affiliation Department of Zoology and Physiology, University of Wyoming, Laramie, United States of America

Mahmoud-Reza Hemami

Roles Conceptualization, Funding acquisition, Investigation, Project administration, Supervision, Writing – review & editing

Affiliation Department of Natural Resources, Isfahan University of Technology, Isfahan, Iran

Jacob R. Goheen

Roles Conceptualization, Methodology, Supervision, Writing – review & editing

Affiliation Department of Zoology and Physiology, University of Wyoming, Laramie, United States of America

Competing Interests

This study was partly funded by Cleveland Metroparks Zoo. There are no patents, products in development or marketed products to declare. This does not alter our adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.