About the Authors

Steven Grant Waldrop

Roles Conceptualization, Data curation, Formal analysis, Investigation, Visualization, Writing – original draft, Writing – review & editing

Affiliation Department of Biomedical Sciences and Pathobiology, Virginia-Maryland College of Veterinary Medicine, Virginia Tech, Blacksburg, Virginia, United States of America

Nammalwar Sriranganathan

Roles Funding acquisition, Resources, Supervision, Validation, Visualization, Writing – review & editing

nathans@vt.edu

Affiliation Department of Biomedical Sciences and Pathobiology, Virginia-Maryland College of Veterinary Medicine, Virginia Tech, Blacksburg, Virginia, United States of America

Competing Interests

We have the following interests. This study was partly funded by Smithfield Foods Murphy Brown, LLC. (Virginia Tech Foundation, Inc. Fund #444115). There are no patents, products in development or marketed products to declare. This does not alter our adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.