About the Authors

Huaihai Chen

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliation Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America

Zamin K. Yang

Roles Formal analysis, Investigation, Methodology

Affiliation Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America

Dan Yip

Roles Investigation, Methodology

Affiliation Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America

Reese H. Morris

Roles Investigation, Methodology

Affiliation Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America

Steven J. Lebreux

Roles Investigation, Methodology

Affiliation Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America

Melissa A. Cregger

Roles Conceptualization, Formal analysis, Funding acquisition, Supervision, Writing – review & editing

Affiliation Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America

Dawn M. Klingeman

Roles Investigation, Methodology, Writing – review & editing

Affiliation Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America

Dafeng Hui

Roles Investigation, Methodology, Writing – review & editing

Affiliation Department of Biological Sciences, Tennessee State University, Nashville, Tennessee, United States of America

Robert L. Hettich

Roles Conceptualization, Funding acquisition, Writing – review & editing

Affiliations Chemical Sciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America, Department of Microbiology, University of Tennessee, Knoxville, Tennessee, United States of America

Steven W. Wilhelm

Roles Conceptualization, Funding acquisition, Writing – review & editing

Affiliation Department of Microbiology, University of Tennessee, Knoxville, Tennessee, United States of America

Gangsheng Wang

Roles Conceptualization, Funding acquisition, Writing – review & editing

Affiliations Environmental Science Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America, Institute for Environmental Genomics and Department of Microbiology & Plant Biology, University of Oklahoma, Norman, Oklahoma, United States of America

Frank E. Löffler

Roles Conceptualization, Funding acquisition, Writing – review & editing

Affiliations Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America, Department of Microbiology, University of Tennessee, Knoxville, Tennessee, United States of America

Christopher W. Schadt

Roles Conceptualization, Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Writing – original draft, Writing – review & editing

schadtcw@ornl.gov

Affiliations Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America, Department of Microbiology, University of Tennessee, Knoxville, Tennessee, United States of America

Competing Interests

The authors of this manuscript have read the journal’s policy and have the following competing interest: DH is an Academic Editor for PLOS ONE. This manuscript has been authored by UT-Battelle, LLC under Contract No. DE-AC05-00OR22725 with the U.S. Department of Energy. The United States Government retains and the publisher, by accepting the article for publication, acknowledges that the United States Government retains a non-exclusive, paid-up, irrevocable, world-wide license to publish or reproduce the published form of this manuscript, or allow others to do so, for United States Government purposes. The Department of Energy will provide public access to these results of federally sponsored research in accordance with the DOE Public Access Plan (http://energy.gov/downloads/doe-public-access-plan). This does not alter our adherence to PLOS ONE policies on sharing data and materials.