About the Authors

Fasil Tadesse Tewolde

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Software, Writing – original draft, Writing – review & editing

Affiliations Graduate School of Horticulture, Chiba University, Matsudo, Japan, Center for Environment, Health and Field Science, Chiba University, Kashiwa, Japan, Ethiopian Institute of Agricultural Research, Addis Ababa, Ethiopia

Kouta Shiina

Roles Writing – review & editing

Affiliation JA Zen-Noh, Kashiwa, Japan

Toru Maruo

Roles Writing – review & editing

Affiliations Graduate School of Horticulture, Chiba University, Matsudo, Japan, Center for Environment, Health and Field Science, Chiba University, Kashiwa, Japan

Michiko Takagaki

Roles Writing – review & editing

Affiliations Graduate School of Horticulture, Chiba University, Matsudo, Japan, Center for Environment, Health and Field Science, Chiba University, Kashiwa, Japan

Toyoki Kozai

Roles Writing – review & editing

Affiliation Japan Plant Factory Association, Kashiwa, Japan

Wataru Yamori

Roles Conceptualization, Supervision, Writing – original draft, Writing – review & editing

wataru.yamori@bs.s.u-tokyo.ac.jp

Affiliations Center for Environment, Health and Field Science, Chiba University, Kashiwa, Japan, Department of Biological Sciences, Faculty of Science, the University of Tokyo, Tokyo, Japan

Competing Interests

The authors declare that the research was conducted in the absence of any commercial or financial relationships that could be construed as a potential conflict of interest. KS is employed by JA Zen-Noh. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.