About the Authors

Xiaolong Hu

Contributed equally to this work with: Xiaolong Hu, Yuting Wei

Roles Data curation, Methodology, Validation, Writing – original draft

Affiliation Laboratory for Non-invasive Research Technology for Endangered Species, College of Nature Conservation, Beijing Forestry University, Beijing, China

Yuting Wei

Contributed equally to this work with: Xiaolong Hu, Yuting Wei

Roles Methodology, Resources, Software

Affiliation Laboratory for Non-invasive Research Technology for Endangered Species, College of Nature Conservation, Beijing Forestry University, Beijing, China

Songlin Huang

Roles Conceptualization, Formal analysis, Investigation, Supervision

Affiliation China Wildlife Mark Center, Chinese Academy of Forestry, Beijing, China

Gang Liu

Roles Data curation, Resources, Software, Writing – review & editing

Affiliation Institute of Wetland Research, Chinese Academy of Forestry, Beijing, China

Yihua Wang

Roles Data curation, Resources, Writing – original draft

Affiliation Laboratory for Non-invasive Research Technology for Endangered Species, College of Nature Conservation, Beijing Forestry University, Beijing, China

Defu Hu

Roles Data curation, Funding acquisition, Project administration, Supervision, Writing – original draft, Writing – review & editing

hudefu2016@163.com (DH); liushuqiang76@sina.com (SL)

Affiliation Laboratory for Non-invasive Research Technology for Endangered Species, College of Nature Conservation, Beijing Forestry University, Beijing, China

Shuqiang Liu

Roles Formal analysis, Funding acquisition, Project administration, Resources

hudefu2016@163.com (DH); liushuqiang76@sina.com (SL)

Affiliation Laboratory for Non-invasive Research Technology for Endangered Species, College of Nature Conservation, Beijing Forestry University, Beijing, China

Competing Interests

There is one commercial funder to declare: Zhangzhou Pien Tze Huang Pharmaceutical Co., Ltd. This company will not use data from this study to develop products, apply for patent, or do any other commercial operation. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials. The authors have no other competing interests to declare.