About the Authors

Olla Al-Jaibaji

Contributed equally to this work with: Olla Al-Jaibaji, Stephen Swioklo

Roles Data curation, Formal analysis, Investigation, Software, Writing – original draft, Writing – review & editing

Affiliation Institute of Genetic Medicine, Newcastle University, Newcastle upon Tyne, United Kingdom

Stephen Swioklo

Contributed equally to this work with: Olla Al-Jaibaji, Stephen Swioklo

Roles Data curation, Writing – review & editing

Affiliation Institute of Genetic Medicine, Newcastle University, Newcastle upon Tyne, United Kingdom

Kristel Gijbels

Roles Resources

‡ These authors also contributed equally to this work.

Affiliation ReGenesys BVBA, Heverlee, Belgium

Bart Vaes

Roles Writing – review & editing

‡ These authors also contributed equally to this work.

Affiliation ReGenesys BVBA, Heverlee, Belgium

Francisco C. Figueiredo

Roles Writing – review & editing

Affiliation Department of Ophthalmology, Royal Victoria Infirmary, Newcastle upon Tyne, United Kingdom

Che J. Connon

Roles Supervision, Writing – review & editing

Che.Connon@newcastle.ac.uk

Affiliation Institute of Genetic Medicine, Newcastle University, Newcastle upon Tyne, United Kingdom

Competing Interests

The authors declare no competing of interest. The commercial affiliation does not alter our adherence to PLOS ONE policies on sharing data and protocols.