About the Authors

Sarah Aileen Lewis

Contributed equally to this work with: Sarah Aileen Lewis, Tetsuya Takimoto

Roles Conceptualization, Data curation, Formal analysis, Investigation, Project administration, Supervision, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliation Department of Medical Genetics, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Tetsuya Takimoto

Contributed equally to this work with: Sarah Aileen Lewis, Tetsuya Takimoto

Roles Conceptualization, Data curation, Formal analysis, Investigation, Project administration, Supervision, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliations Department of Medical Genetics, University of Wisconsin-Madison, Madison, Wisconsin, United States of America, Institute for Innovation, Ajinomoto Co., Inc., Tokyo, Japan

Shima Mehrvar

Roles Data curation, Formal analysis, Investigation, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliation Department of Electrical Engineering, Biophotonics Laboratory, University of Wisconsin, Milwaukee, Wisconsin, United States of America

Hitoshi Higuchi

Roles Conceptualization, Data curation, Investigation, Methodology, Validation, Writing – review & editing

Current address: Department of Dental Anesthesiology, Okayama University Hospital, Okayama, Japan

Affiliation Department of Medical Genetics, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Anna-Lisa Doebley

Roles Investigation, Validation

Current address: Molecular and Cellular Biology Program, University of Washington, Seattle, Washington, United States of America

Affiliation Department of Medical Genetics, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Giangela Stokes

Roles Investigation, Validation

Affiliation Department of Medical Genetics, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Nader Sheibani

Roles Conceptualization, Funding acquisition, Methodology, Project administration, Writing – original draft, Writing – review & editing

Affiliations Department Ophthalmology and Visual Sciences, Biomedical Engineering, and Cell and Regenerative Biology, University of Wisconsin-Madison, Madison, Wisconsin, United States of America, McPherson Eye Research Institute, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Sakae Ikeda

Roles Conceptualization, Methodology, Resources, Supervision, Writing – original draft, Writing – review & editing

Affiliations Department of Medical Genetics, University of Wisconsin-Madison, Madison, Wisconsin, United States of America, McPherson Eye Research Institute, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Mahsa Ranji

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliation Department of Electrical Engineering, Biophotonics Laboratory, University of Wisconsin, Milwaukee, Wisconsin, United States of America

Akihiro Ikeda

Roles Conceptualization, Funding acquisition, Methodology, Project administration, Resources, Supervision, Writing – original draft, Writing – review & editing

aikeda@wisc.edu

Affiliations Department of Medical Genetics, University of Wisconsin-Madison, Madison, Wisconsin, United States of America, McPherson Eye Research Institute, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Competing Interests

I have read the journal’s policy and the authors of this manuscript have the following competing interests. TT was employed by Ajinomoto Co., Inc., while working on this project as a Visiting Scholar at University of Wisconsin-Madison. This does not alter our adherence to PLOS ONE policies on sharing data and materials.