About the Authors

David M. Brown

Roles Conceptualization, Formal analysis, Investigation, Writing – original draft

Affiliation School of Biosciences, University of Nottingham, Sutton Bonington Campus, Loughborough, United Kingdom

Sarah Jones

Roles Formal analysis, Investigation

Affiliation School of Biosciences, University of Nottingham, Sutton Bonington Campus, Loughborough, United Kingdom

Zoe C. T. R. Daniel

Roles Formal analysis, Investigation

Affiliation School of Biosciences, University of Nottingham, Sutton Bonington Campus, Loughborough, United Kingdom

Madelaine C. Brearley

Roles Formal analysis, Methodology

Affiliation School of Biosciences, University of Nottingham, Sutton Bonington Campus, Loughborough, United Kingdom

Jo E. Lewis

Roles Data curation, Formal analysis

Affiliation School of Life Sciences, University of Nottingham, Medical School, Nottingham, United Kingdom

Francis J. P. Ebling

Roles Conceptualization, Funding acquisition, Supervision, Writing – review & editing

Affiliation School of Life Sciences, University of Nottingham, Medical School, Nottingham, United Kingdom

Tim Parr

Roles Conceptualization, Funding acquisition, Supervision, Writing – review & editing

Affiliation School of Biosciences, University of Nottingham, Sutton Bonington Campus, Loughborough, United Kingdom

John M. Brameld

Roles Conceptualization, Funding acquisition, Supervision, Writing – review & editing

John.Brameld@Nottingham.ac.uk

Affiliation School of Biosciences, University of Nottingham, Sutton Bonington Campus, Loughborough, United Kingdom

Competing Interests

This research was funded by the BBSRC and Zoetis (formally Pfizer Animal Health). The funders were not involved in the study design, collection, analysis or interpretation of the data, nor in the writing of the paper. Zoetis approved the decision to submit for publication. This does not alter our adherence to PLOS ONE policies on sharing data and materials.