About the Authors

Yi-Chen Chen

Contributed equally to this work with: Yi-Chen Chen, Wei-Che Chang

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Validation, Writing – original draft

Affiliations Institute of Biotechnology, National Taiwan University, Taipei, Taiwan, Univ Lyon, Université Lyon 1, INSERM, INRA, Stem Cell and Brain Research Institute, U1208, USC1361, Bron, France

Wei-Che Chang

Contributed equally to this work with: Yi-Chen Chen, Wei-Che Chang

Roles Conceptualization, Formal analysis, Investigation, Methodology

Affiliation Department of Animal Science and Technology, National Taiwan University, Taipei, Taiwan

Shau-Ping Lin

Roles Conceptualization, Data curation, Formal analysis, Supervision, Validation, Writing – review & editing

Affiliations Institute of Biotechnology, National Taiwan University, Taipei, Taiwan, Agricultural Biotechnology Research Center, Academia Sinica, Taipei, Taiwan

Masataka Minami

Roles Methodology, Resources

Affiliation Nissan Chemical Corporation, Tokyo, Japan

Christian Jean

Roles Data curation, Methodology

Affiliation Univ Lyon, Université Lyon 1, INSERM, INRA, Stem Cell and Brain Research Institute, U1208, USC1361, Bron, France

Hisato Hayashi

Roles Methodology, Resources

Affiliation Nissan Chemical Corporation, Tokyo, Japan

Sylvie Rival-Gervier

Roles Conceptualization, Investigation, Methodology

Affiliation Univ Lyon, Université Lyon 1, INSERM, INRA, Stem Cell and Brain Research Institute, U1208, USC1361, Bron, France

Tatsuro Kanaki

Roles Methodology, Resources

Affiliation Nissan Chemical Corporation, Tokyo, Japan

Shinn-Chih Wu

Roles Funding acquisition, Supervision, Writing – review & editing

scw01@ntu.edu.tw (SCW); bertrand.pain@inserm.fr (BP)

Affiliations Institute of Biotechnology, National Taiwan University, Taipei, Taiwan, Department of Animal Science and Technology, National Taiwan University, Taipei, Taiwan

Bertrand Pain

Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Supervision, Validation, Writing – original draft, Writing – review & editing

scw01@ntu.edu.tw (SCW); bertrand.pain@inserm.fr (BP)

Affiliation Univ Lyon, Université Lyon 1, INSERM, INRA, Stem Cell and Brain Research Institute, U1208, USC1361, Bron, France

Competing Interests

Nissan Chemical Corporation provided support in the form of salaries for authors MM, HH, and TK, but did not have any additional role in the study design, data collection and analysis, decision to publish, or preparation of the manuscript. The specific roles of these authors are articulated in the ‘author contributions’ section. This does not alter our adherence to PLOS ONE policies on sharing data and materials.