About the Authors

Tomoyoshi Inoue

Contributed equally to this work with: Tomoyoshi Inoue, Yoshiya Ito

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliations Department of Molecular Pharmacology, Graduate School of Medical Sciences, Kitasato University, Sagamihara, Kanagawa, Japan, Department of Gastroenterology, Kitasato University School of Medicine, Sagamihara, Kanagawa, Japan, Department of Pharmacology, Kitasato University School of Medicine, Sagamihara, Kanagawa, Japan

Yoshiya Ito

Contributed equally to this work with: Tomoyoshi Inoue, Yoshiya Ito

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliations Department of Molecular Pharmacology, Graduate School of Medical Sciences, Kitasato University, Sagamihara, Kanagawa, Japan, Department of Pharmacology, Kitasato University School of Medicine, Sagamihara, Kanagawa, Japan

Nobuyuki Nishizawa

Roles Data curation, Formal analysis, Investigation, Methodology

Affiliations Department of Molecular Pharmacology, Graduate School of Medical Sciences, Kitasato University, Sagamihara, Kanagawa, Japan, Department of Surgery, Kitasato University School of Medicine, Sagamihara, Kanagawa, Japan

Koji Eshima

Roles Data curation, Formal analysis, Investigation, Methodology, Supervision, Validation

Affiliation Department of Immunology, Kitasato University School of Medicine, Sagamihara, Kanagawa, Japan

Ken Kojo

Roles Data curation, Formal analysis, Investigation, Methodology

Affiliation Department of Immunology, Kitasato University School of Medicine, Sagamihara, Kanagawa, Japan

Fumisato Otaka

Roles Data curation, Formal analysis, Investigation, Methodology

Affiliations Department of Molecular Pharmacology, Graduate School of Medical Sciences, Kitasato University, Sagamihara, Kanagawa, Japan, Department of Gastroenterology, Kitasato University School of Medicine, Sagamihara, Kanagawa, Japan, Department of Pharmacology, Kitasato University School of Medicine, Sagamihara, Kanagawa, Japan

Tomohiro Betto

Roles Data curation, Formal analysis, Investigation, Methodology

Affiliations Department of Molecular Pharmacology, Graduate School of Medical Sciences, Kitasato University, Sagamihara, Kanagawa, Japan, Department of Gastroenterology, Kitasato University School of Medicine, Sagamihara, Kanagawa, Japan, Department of Pharmacology, Kitasato University School of Medicine, Sagamihara, Kanagawa, Japan

Sakiko Yamane

Roles Data curation, Formal analysis, Investigation, Methodology

Affiliations Department of Molecular Pharmacology, Graduate School of Medical Sciences, Kitasato University, Sagamihara, Kanagawa, Japan, Department of Gastroenterology, Kitasato University School of Medicine, Sagamihara, Kanagawa, Japan, Department of Pharmacology, Kitasato University School of Medicine, Sagamihara, Kanagawa, Japan

Kazutake Tsujikawa

Roles Methodology, Resources

Affiliation Laboratory of Molecular and Cellular Physiology, Graduate School of Pharmaceutical Sciences, Osaka University, Osaka, Japan

Wasaburo Koizumi

Roles Funding acquisition, Project administration

Affiliation Department of Gastroenterology, Kitasato University School of Medicine, Sagamihara, Kanagawa, Japan

Masataka Majima

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Project administration, Resources, Supervision, Validation, Writing – original draft, Writing – review & editing

mmajima@med.kitasto-u.ac.jp

Affiliations Department of Molecular Pharmacology, Graduate School of Medical Sciences, Kitasato University, Sagamihara, Kanagawa, Japan, Department of Pharmacology, Kitasato University School of Medicine, Sagamihara, Kanagawa, Japan

Competing Interests

The authors have declared that no competing interests exist.