About the Authors

He Zhou

Contributed equally to this work with: He Zhou, Qi-Zheng Xu

Roles Investigation

Affiliation Key Laboratory of Mariculture and Stock Enhancement in North China’s Sea, Ministry of Agriculture, Dalian Ocean University, Dalian, China

Qi-Zheng Xu

Contributed equally to this work with: He Zhou, Qi-Zheng Xu

Roles Formal analysis, Investigation, Writing – original draft

Affiliation Key Laboratory of Mariculture and Stock Enhancement in North China’s Sea, Ministry of Agriculture, Dalian Ocean University, Dalian, China

Rui Zhang

Roles Investigation

Affiliation Key Laboratory of Mariculture and Stock Enhancement in North China’s Sea, Ministry of Agriculture, Dalian Ocean University, Dalian, China

Zi-Xin Zhuang

Roles Investigation

Affiliation Key Laboratory of Mariculture and Stock Enhancement in North China’s Sea, Ministry of Agriculture, Dalian Ocean University, Dalian, China

Yin-Qiang Ma

Roles Data curation

Affiliation Dalian Medical University, Dalian, China

Wei Wang

Roles Investigation

Affiliation Key Laboratory of Mariculture and Stock Enhancement in North China’s Sea, Ministry of Agriculture, Dalian Ocean University, Dalian, China

Tian-Yu Ma

Roles Investigation

Affiliation Key Laboratory of Mariculture and Stock Enhancement in North China’s Sea, Ministry of Agriculture, Dalian Ocean University, Dalian, China

Yi Sui

Roles Investigation

Affiliation Ma'anshan Municipal Agriculture Commission, Anhui, China

Yang Liu

Roles Data curation

Affiliation Key Laboratory of Mariculture and Stock Enhancement in North China’s Sea, Ministry of Agriculture, Dalian Ocean University, Dalian, China

Xiaojuan Cao

Roles Project administration, Writing – review & editing

caoxiaojuan@mail.hzau.edu.cn

Affiliation College of Fisheries, Key Lab of Agricultural Animal Genetics, Breeding and Reproduction of Ministry of Education/Key Lab of Freshwater Animal Breeding, Ministry of Agriculture, Huazhong Agricultural University, Wuhan, Hubei, China

Competing Interests

The authors have declared that no competing interests exist.