About the Authors

Sarah Bouzroud

Contributed equally to this work with: Sarah Bouzroud, Sandra Gouiaa, Nan Hu

Roles Formal analysis, Investigation, Methodology, Writing – original draft

Affiliations GBF, Université de Toulouse, INRA, Castanet-Tolosan, France, Laboratoire de physiologie et biotechnologie Végétales, Centre de biotechnologie végétale et microbienne biodiversité et environnement, Faculté des Sciences, Université Mohammed V de Rabat, Rabat, Maroc

Sandra Gouiaa

Contributed equally to this work with: Sarah Bouzroud, Sandra Gouiaa, Nan Hu

Roles Formal analysis, Investigation, Methodology

Affiliation GBF, Université de Toulouse, INRA, Castanet-Tolosan, France

Nan Hu

Contributed equally to this work with: Sarah Bouzroud, Sandra Gouiaa, Nan Hu

Roles Formal analysis, Investigation, Methodology

Affiliation GBF, Université de Toulouse, INRA, Castanet-Tolosan, France

Anne Bernadac

Roles Conceptualization, Supervision, Validation, Writing – original draft, Writing – review & editing

Affiliation GBF, Université de Toulouse, INRA, Castanet-Tolosan, France

Isabelle Mila

Roles Methodology, Resources, Writing – review & editing

Affiliation GBF, Université de Toulouse, INRA, Castanet-Tolosan, France

Najib Bendaou

Roles Conceptualization, Writing – review & editing

Affiliation Laboratoire de physiologie et biotechnologie Végétales, Centre de biotechnologie végétale et microbienne biodiversité et environnement, Faculté des Sciences, Université Mohammed V de Rabat, Rabat, Maroc

AbdelAziz Smouni

Roles Conceptualization, Supervision, Validation, Writing – original draft, Writing – review & editing

Affiliation Laboratoire de physiologie et biotechnologie Végétales, Centre de biotechnologie végétale et microbienne biodiversité et environnement, Faculté des Sciences, Université Mohammed V de Rabat, Rabat, Maroc

Mondher Bouzayen

Roles Conceptualization, Supervision, Validation, Writing – original draft, Writing – review & editing

Affiliation GBF, Université de Toulouse, INRA, Castanet-Tolosan, France

Mohamed Zouine

Roles Conceptualization, Supervision, Validation, Writing – original draft, Writing – review & editing

mohamed.zouine@ensat.fr

Affiliation GBF, Université de Toulouse, INRA, Castanet-Tolosan, France

Competing Interests

All authors have read, edited, and approved the final version of the manuscript. They have declared that no competing interests exist.