About the Authors

Ruairidh D. Campbell

Contributed equally to this work with: Ruairidh D. Campbell, Frank Rosell

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Writing – original draft, Writing – review & editing

Current address: Scottish Natural Heritage, Great Glen House, Inverness, United Kingdom

‡ CN and DWM also contributed equally to this work. RDC and FR are joint first authors on this work.

Affiliations University College of Southeast Norway, Department of Natural Sciences and Environmental Health, Bø i Telemark, Norway, Wildlife Conservation Research Unit, The Recanati-Kaplan Centre, Zoology Department, University of Oxford, Tubney, United Kingdom

Frank Rosell

Contributed equally to this work with: Ruairidh D. Campbell, Frank Rosell

Roles Data curation, Funding acquisition, Methodology, Project administration, Resources, Supervision, Writing – original draft

frank.rosell@usn.no

‡ CN and DWM also contributed equally to this work. RDC and FR are joint first authors on this work.

Affiliation University College of Southeast Norway, Department of Natural Sciences and Environmental Health, Bø i Telemark, Norway

Chris Newman

Roles Writing – original draft, Writing – review & editing

‡ CN and DWM also contributed equally to this work. RDC and FR are joint first authors on this work.

Affiliation Wildlife Conservation Research Unit, The Recanati-Kaplan Centre, Zoology Department, University of Oxford, Tubney, United Kingdom

David W. Macdonald

Roles Funding acquisition, Supervision, Writing – original draft

‡ CN and DWM also contributed equally to this work. RDC and FR are joint first authors on this work.

Affiliation Wildlife Conservation Research Unit, The Recanati-Kaplan Centre, Zoology Department, University of Oxford, Tubney, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.