About the Authors

Ying-Han Wang

Roles Conceptualization, Investigation, Methodology

Affiliations Department of Life Science, National Taiwan Normal University, Taipei, Taiwan, Institute of Ecology and Evolutionary Biology, National Taiwan University, Taipei, Taiwan

Yu-Wei Hsiao

Contributed equally to this work with: Yu-Wei Hsiao, Ko-Huan Lee

Roles Conceptualization, Investigation, Methodology

Affiliation Department of Life Science, National Taiwan Normal University, Taipei, Taiwan

Ko-Huan Lee

Contributed equally to this work with: Yu-Wei Hsiao, Ko-Huan Lee

Roles Formal analysis, Investigation, Writing – original draft

Affiliation Department of Life Science, National Taiwan Normal University, Taipei, Taiwan

Hui-Yun Tseng

Roles Data curation, Resources

Affiliation Department of Biology, National Museum of Natural Science, Taichung, Taiwan

Yen-Po Lin

Roles Data curation, Resources

Affiliation Division of Zoology, Taiwan Endemic Species Research Institute, Nantou, Taiwan

Shohei Komaki

Roles Resources, Supervision, Writing – review & editing

cynops00pyrrhogaster@gmail.com (SK); lizard.dna@gmail.com (SML)

Affiliations Division of Developmental Science, Graduate School for International Development and Cooperation, Hiroshima University, Hiroshima, Japan, Global Career Design Center, Hiroshima University, Hiroshima, Japan

Si-Min Lin

Roles Conceptualization, Project administration, Supervision, Writing – original draft, Writing – review & editing

cynops00pyrrhogaster@gmail.com (SK); lizard.dna@gmail.com (SML)

Affiliation Department of Life Science, National Taiwan Normal University, Taipei, Taiwan

Competing Interests

The authors have declared that no competing interests exist.