About the Authors

Alan M. Watson

Roles Formal analysis, Investigation, Methodology, Visualization, Writing – original draft, Writing – review & editing

Affiliations Center for Vaccine Research, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Microbiology and Molecular Genetics, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Annika H. Rose

Roles Data curation, Formal analysis

Affiliation Department of Neurobiology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Gregory A. Gibson

Roles Data curation

Affiliation Center for Biologic Imaging and the Department of Cellular Biology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Christina L. Gardner

Roles Data curation

Affiliations Center for Vaccine Research, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Microbiology and Molecular Genetics, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Chengqun Sun

Roles Resources

Affiliations Center for Vaccine Research, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Microbiology and Molecular Genetics, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Douglas S. Reed

Roles Resources

Affiliations Center for Vaccine Research, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Immunology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

L. K. Metthew Lam

Roles Data curation

Affiliations Center for Vaccine Research, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Microbiology and Molecular Genetics, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Claudette M. St. Croix

Roles Writing – review & editing

Affiliation Center for Biologic Imaging and the Department of Cellular Biology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Peter L. Strick

Roles Funding acquisition, Resources, Writing – review & editing

Affiliation Department of Neurobiology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

William B. Klimstra

Roles Conceptualization, Funding acquisition, Resources, Supervision, Writing – review & editing

simon.watkins@pitt.edu (SCW); klimstra@pitt.edu (WBK)

Affiliations Center for Vaccine Research, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Microbiology and Molecular Genetics, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Simon C. Watkins

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Methodology, Resources, Supervision, Visualization, Writing – original draft, Writing – review & editing

simon.watkins@pitt.edu (SCW); klimstra@pitt.edu (WBK)

Affiliation Center for Biologic Imaging and the Department of Cellular Biology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.