About the Authors

Youngjune Bhak

Affiliations The Genomics Institute, Ulsan National Institute of Science and Technology (UNIST), Ulsan, Republic of Korea, Department of Biomedical Engineering, School of Life Sciences, Ulsan National Institute of Science and Technology (UNIST), Ulsan, Republic of Korea

Yeonsu Jeon

Affiliations The Genomics Institute, Ulsan National Institute of Science and Technology (UNIST), Ulsan, Republic of Korea, Department of Biomedical Engineering, School of Life Sciences, Ulsan National Institute of Science and Technology (UNIST), Ulsan, Republic of Korea

Sungwon Jeon

Affiliations The Genomics Institute, Ulsan National Institute of Science and Technology (UNIST), Ulsan, Republic of Korea, Department of Biomedical Engineering, School of Life Sciences, Ulsan National Institute of Science and Technology (UNIST), Ulsan, Republic of Korea

Oksung Chung

Affiliations Personal Genomics Institute, Genome Research Foundation, Cheongju, Republic of Korea, Geromics, Ulsan, Republic of Korea

Sungwoong Jho

Affiliation Personal Genomics Institute, Genome Research Foundation, Cheongju, Republic of Korea

JeHoon Jun

Affiliations Personal Genomics Institute, Genome Research Foundation, Cheongju, Republic of Korea, Geromics, Ulsan, Republic of Korea

Hak-Min Kim

Affiliations The Genomics Institute, Ulsan National Institute of Science and Technology (UNIST), Ulsan, Republic of Korea, Department of Biomedical Engineering, School of Life Sciences, Ulsan National Institute of Science and Technology (UNIST), Ulsan, Republic of Korea

Yongsoo Cho

Affiliations The Genomics Institute, Ulsan National Institute of Science and Technology (UNIST), Ulsan, Republic of Korea, Department of Biomedical Engineering, School of Life Sciences, Ulsan National Institute of Science and Technology (UNIST), Ulsan, Republic of Korea

Changhan Yoon

Affiliations The Genomics Institute, Ulsan National Institute of Science and Technology (UNIST), Ulsan, Republic of Korea, Department of Biomedical Science, School of Nano-Bioscience & chemical Engineering, Ulsan National Institute of Science and Technology (UNIST), Ulsan, Republic of Korea

Seungwoo Lee

Affiliation BK21 PLUS Program for Creative Veterinary Science Research, Research Institute for Veterinary Science, and College of Veterinary Medicine, Seoul National University, Seoul, Republic of Korea

Jung-Hoon Kang

Affiliation National Research Institute of Cultural Heritage, Cultural Heritage Administration, Daejeon, Republic of Korea

Jong-Deock Lim

Affiliation National Research Institute of Cultural Heritage, Cultural Heritage Administration, Daejeon, Republic of Korea

Junghwa An

Affiliation Animal Resources Division, National Institute of Biological Resources, Incheon, Republic of Korea

Yun Sung Cho

joys0406@gmail.com (YSC); dyryu@snu.ac.kr (DYR); jongbhak@genomics.org (JB)

Affiliations The Genomics Institute, Ulsan National Institute of Science and Technology (UNIST), Ulsan, Republic of Korea, Department of Biomedical Engineering, School of Life Sciences, Ulsan National Institute of Science and Technology (UNIST), Ulsan, Republic of Korea, Personal Genomics Institute, Genome Research Foundation, Cheongju, Republic of Korea

Doug-Young Ryu

joys0406@gmail.com (YSC); dyryu@snu.ac.kr (DYR); jongbhak@genomics.org (JB)

Affiliation BK21 PLUS Program for Creative Veterinary Science Research, Research Institute for Veterinary Science, and College of Veterinary Medicine, Seoul National University, Seoul, Republic of Korea

Jong Bhak

joys0406@gmail.com (YSC); dyryu@snu.ac.kr (DYR); jongbhak@genomics.org (JB)

Affiliations The Genomics Institute, Ulsan National Institute of Science and Technology (UNIST), Ulsan, Republic of Korea, Department of Biomedical Engineering, School of Life Sciences, Ulsan National Institute of Science and Technology (UNIST), Ulsan, Republic of Korea, Personal Genomics Institute, Genome Research Foundation, Cheongju, Republic of Korea, Geromics, Ulsan, Republic of Korea

Competing Interests

OC and JHJ are employees and JB is a founder of Geromics Inc. This does not alter our adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceptualization: YB YSC DYR JB. Data curation: YB OC S. Jho JHJ HMK. Formal analysis: YB YJ S. Jeon. Funding acquisition: JB. Investigation: YB YJ S. Jeon YC. Methodology: JB YB. Project administration: JA DYR JB. Resources: DYR SL JHK JDL. Software: YB YJ S. Jeon OC S. Jho JHJ HMK. Supervision: JB. Validation: YB CY. Visualization: YB S. Jeon. Writing – original draft: JB YB. Writing – review & editing: YB YJ S. Jeon JA YSC JB.