About the Authors

Tingting Zhao

Contributed equally to this work with: Tingting Zhao, Zemin Wang

Affiliations Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan, P.R. China, University of Chinese Academy of Sciences, Beijing, P.R. China

Zemin Wang

Contributed equally to this work with: Tingting Zhao, Zemin Wang

Affiliation Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan, P.R. China

Lingye Su

Affiliations University of Chinese Academy of Sciences, Beijing, P.R. China, Beijing Key Laboratory of Grape Sciences and Enology, CAS Key Laboratory of Plant Resources, Institute of Botany, Chinese Academy of Sciences, Beijing, P.R. China

Xiaoming Sun

Affiliations Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan, P.R. China, Beijing Key Laboratory of Grape Sciences and Enology, CAS Key Laboratory of Plant Resources, Institute of Botany, Chinese Academy of Sciences, Beijing, P.R. China

Jun Cheng

Affiliation Beijing Key Laboratory of Grape Sciences and Enology, CAS Key Laboratory of Plant Resources, Institute of Botany, Chinese Academy of Sciences, Beijing, P.R. China

Langlang Zhang

Affiliations Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan, P.R. China, University of Chinese Academy of Sciences, Beijing, P.R. China

Sospeter Karanja Karungo

Affiliations Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan, P.R. China, University of Chinese Academy of Sciences, Beijing, P.R. China

Yuepeng Han

Affiliation Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan, P.R. China

Shaohua Li

Affiliations Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan, P.R. China, Beijing Key Laboratory of Grape Sciences and Enology, CAS Key Laboratory of Plant Resources, Institute of Botany, Chinese Academy of Sciences, Beijing, P.R. China

Haiping Xin

xinhaiping215@hotmail.com

Affiliation Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan, P.R. China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: HX SL ZW TZ. Formal analysis: TZ ZW. Funding acquisition: HX. Investigation: TZ ZW LS LZ XS JC SK. Methodology: HX ZW XS JC. Project administration: HX. Resources: YH. Supervision: HX. Visualization: TZ ZW. Writing – original draft: TZ ZW. Writing – review & editing: HX.