About the Authors

Yulong Duan

Contributed equally to this work with: Yulong Duan, Fasi Wu

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Software, Visualization, Writing – original draft

‡ These authors are co-first authors on this work.

Affiliation Key Laboratory of Extreme Environmental Microbial Resources and Engineering, Gansu Province, Northwest Institute of Eco-Environment and Resources, University of Chinese Academy of Sciences, Lanzhou, P.R.China

Fasi Wu

Contributed equally to this work with: Yulong Duan, Fasi Wu

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Resources, Software, Supervision, Validation, Visualization, Writing – original draft

‡ These authors are co-first authors on this work.

Affiliations National Research Center for Conservation of Ancient Wall Paintings and Earthen Sites, Dunhuang Academy, Dunhuang, Gansu, P.R.China, MOE Key Laboratory of Cell Activities and Stress Adaptations, School of Life Sciences, Lanzhou University, Lanzhou, P.R.China

Wanfu Wang

Roles Conceptualization, Funding acquisition, Investigation, Project administration, Resources, Supervision, Validation, Visualization, Writing – original draft

wwanfu@hotmail.com (WFW); fenghy@lzu.edu.cn (HYF)

Affiliations Key Laboratory of Extreme Environmental Microbial Resources and Engineering, Gansu Province, Northwest Institute of Eco-Environment and Resources, University of Chinese Academy of Sciences, Lanzhou, P.R.China, National Research Center for Conservation of Ancient Wall Paintings and Earthen Sites, Dunhuang Academy, Dunhuang, Gansu, P.R.China, MOE Key Laboratory of Cell Activities and Stress Adaptations, School of Life Sciences, Lanzhou University, Lanzhou, P.R.China

Dongpeng He

Roles Investigation

Affiliation National Research Center for Conservation of Ancient Wall Paintings and Earthen Sites, Dunhuang Academy, Dunhuang, Gansu, P.R.China

Ji-Dong Gu

Roles Conceptualization, Writing – review & editing

Affiliation Laboratory of Environmental Microbiology and Toxicology, School of Biological Sciences, The University of Hong Kong, Hong Kong SAR, P.R.China

Huyuan Feng

Roles Conceptualization, Writing – review & editing

wwanfu@hotmail.com (WFW); fenghy@lzu.edu.cn (HYF)

Affiliation MOE Key Laboratory of Cell Activities and Stress Adaptations, School of Life Sciences, Lanzhou University, Lanzhou, P.R.China

Tuo Chen

Roles Conceptualization, Writing – review & editing

Affiliation Key Laboratory of Extreme Environmental Microbial Resources and Engineering, Gansu Province, Northwest Institute of Eco-Environment and Resources, University of Chinese Academy of Sciences, Lanzhou, P.R.China

Guangxiu Liu

Roles Conceptualization, Writing – review & editing

Affiliation Key Laboratory of Extreme Environmental Microbial Resources and Engineering, Gansu Province, Northwest Institute of Eco-Environment and Resources, University of Chinese Academy of Sciences, Lanzhou, P.R.China

Lizhe An

Roles Conceptualization, Writing – review & editing

Affiliations Key Laboratory of Extreme Environmental Microbial Resources and Engineering, Gansu Province, Northwest Institute of Eco-Environment and Resources, University of Chinese Academy of Sciences, Lanzhou, P.R.China, MOE Key Laboratory of Cell Activities and Stress Adaptations, School of Life Sciences, Lanzhou University, Lanzhou, P.R.China

Competing Interests

The authors have declared that no competing interests exist.