About the Authors

Tsutomu Iwata

Affiliations Department of Cell Biology, Graduate School of Medical Sciences, Nagoya City University, Nagoya, Aichi, Japan, Department of Gastroenterological Surgery, Aichi Medical University School of Medicine, Nagakute, Japan

Tairin Uchino

Affiliations Department of Cell Biology, Graduate School of Medical Sciences, Nagoya City University, Nagoya, Aichi, Japan, Department of Gastroenterological Surgery, Aichi Medical University School of Medicine, Nagakute, Japan

Ayako Koyama

Affiliation Department of Cell Biology, Graduate School of Medical Sciences, Nagoya City University, Nagoya, Aichi, Japan

Yoshikazu Johmura

Affiliations Department of Cell Biology, Graduate School of Medical Sciences, Nagoya City University, Nagoya, Aichi, Japan, Division of Cancer Cell Biology, Department of Cancer Biology, Institute of Medical Science, The University of Tokyo, Minato-ku, Tokyo, Japan

Kenichi Koyama

Affiliation Department of Gastroenterological Surgery, Aichi Medical University School of Medicine, Nagakute, Japan

Takuya Saito

Affiliation Department of Gastroenterological Surgery, Aichi Medical University School of Medicine, Nagakute, Japan

Seiji Ishiguro

Affiliation Department of Gastroenterological Surgery, Aichi Medical University School of Medicine, Nagakute, Japan

Takashi Arikawa

Affiliation Department of Gastroenterological Surgery, Aichi Medical University School of Medicine, Nagakute, Japan

Shunichiro Komatsu

Affiliation Department of Gastroenterological Surgery, Aichi Medical University School of Medicine, Nagakute, Japan

Masahiko Miyachi

Affiliation Department of Gastroenterological Surgery, Aichi Medical University School of Medicine, Nagakute, Japan

Tsuyoshi Sano

Affiliation Department of Gastroenterological Surgery, Aichi Medical University School of Medicine, Nagakute, Japan

Makoto Nakanishi

Affiliations Department of Cell Biology, Graduate School of Medical Sciences, Nagoya City University, Nagoya, Aichi, Japan, Division of Cancer Cell Biology, Department of Cancer Biology, Institute of Medical Science, The University of Tokyo, Minato-ku, Tokyo, Japan

Midori Shimada

shimada@yamaguchi-u.ac.jp

Current address: Department of Biochemistry, Joint Faculty of Veterinary Science, Yamaguchi University, Yoshida, Yamaguchi, Japan

Affiliation Department of Cell Biology, Graduate School of Medical Sciences, Nagoya City University, Nagoya, Aichi, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: MN MS. Data curation: TI MS. Formal analysis: TI MN MS. Funding acquisition: MN MS. Investigation: TI TU AK YJ KK T. Saito SI TA SK MM T. Sano. Methodology: TI TU AK YJ KK T. Saito SI TA SK MM T. Sano. Project administration: TI MN MS. Resources: TI MN MS. Software: TI MN MS. Supervision: MN MS. Validation: TI MN MS. Visualization: TI MN MS. Writing – original draft: TI MN MS. Writing – review & editing: TI MN MS.