About the Authors

Catherine Hobaiter

clh42@st-andrews.ac.uk

Affiliations Centre for Social Learning and Cognitive Evolution and Scottish Primate Research Group, School of Psychology and Neuroscience, University of St Andrews, St Andrews, Scotland, Budongo Conservation Field Station, Masindi, Uganda

Liran Samuni

Affiliations Budongo Conservation Field Station, Masindi, Uganda, Department of Primatology, Max Planck Institute for Evolutionary Anthropology, Leipzig, Germany

Caroline Mullins

Affiliations Centre for Social Learning and Cognitive Evolution and Scottish Primate Research Group, School of Psychology and Neuroscience, University of St Andrews, St Andrews, Scotland, Budongo Conservation Field Station, Masindi, Uganda

Walter John Akankwasa

Affiliations Budongo Conservation Field Station, Masindi, Uganda, School of Veterinary Medicine and Animal Resources, Makerere University, Kampala, Uganda

Klaus Zuberbühler

Affiliations Centre for Social Learning and Cognitive Evolution and Scottish Primate Research Group, School of Psychology and Neuroscience, University of St Andrews, St Andrews, Scotland, Budongo Conservation Field Station, Masindi, Uganda, Department of Comparative Cognition, Institute of Biology, University of Neuchatel, Neuchâtel, Switzerland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: CH LS CM WJA KZ. Data curation: CH LS CM WJA. Formal analysis: CH LS CM WJA. Funding acquisition: CH KZ. Investigation: CH LS CM WJA. Methodology: CH LS CM WJA KZ. Project administration: CH LS CM WJA KZ. Resources: CH LS CM WJA KZ. Supervision: CH KZ. Validation: CH LS CM WJA KZ. Writing – original draft: CH LS CM WJA KZ. Writing – review & editing: CH LS CM WJA KZ.