About the Authors

Zhiqiang Wu

Contributed equally to this work with: Zhiqiang Wu, Cuihua Gu

wu.zhiqiang.1020@gmail.com

Affiliation State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing, China

Cuihua Gu

Contributed equally to this work with: Zhiqiang Wu, Cuihua Gu

Affiliation School of Landscape and Architecture, Zhejiang Agriculture and Forestry University, Hangzhou, China

Luke R. Tembrock

Affiliation Department of Biology, Colorado State University, Fort Collins, Colorado, United States of America

Dong Zhang

Affiliation Department of Statistics, Iowa State University, Ames, Iowa, United States of America

Song Ge

Affiliation State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing, China

Competing Interests

The authors report no conflicts of interest. The authors alone are responsible for the content and writing of this article.

Author Contributions

Conceptualization: ZQW SG. Data curation: ZQW CHG SG. Formal analysis: ZQW CHG LRT DZ SG. Funding acquisition: ZQW CHG SG. Investigation: ZQW CHG SG. Methodology: ZQW CHG LRT DZ SG. Project administration: ZQW SG. Resources: ZQW SG. Software: ZQW CHG LRT SG. Supervision: ZQW CHG SG. Validation: ZQW CHG LRT DZ SG. Visualization: ZQW CHG LRT DZ SG. Writing – original draft: ZQW CHG LRT DZ SG. Writing – review & editing: ZQW CHG LRT DZ SG.