About the Authors

Song Zhang

Affiliations State Key Laboratory of Genetic Engineering, Collaborative Innovation Center for Genetics and Development, School of Life Sciences, Fudan University, Shanghai, China, Ministry of Education Key Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan University, Shanghai, China, Center for genetic Epidemiology, School of Life Sciences, Fudan University, Shanghai, China

Qianyi Xiao

Affiliation Center for Genetic Translational Medicine and Prevention, School of Public Health, Fudan University, Shanghai, China

Zhuqing Shi

Affiliations State Key Laboratory of Genetic Engineering, Collaborative Innovation Center for Genetics and Development, School of Life Sciences, Fudan University, Shanghai, China, Ministry of Education Key Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan University, Shanghai, China, Center for genetic Epidemiology, School of Life Sciences, Fudan University, Shanghai, China

Guopeng Yu

Affiliation Department of Urology, The 9th People’s Hospital of Shanghai, School of Medicine, Shanghai Jiaotong University, Shanghai, China

Xiao-Pin Ma

Affiliation State Key Laboratory of Genetic Engineering, Collaborative Innovation Center for Genetics and Development, School of Life Sciences, Fudan University, Shanghai, China

Haitao Chen

Affiliation Center for Genetic Translational Medicine and Prevention, School of Public Health, Fudan University, Shanghai, China

Pengyin Zhang

Affiliations State Key Laboratory of Genetic Engineering, Collaborative Innovation Center for Genetics and Development, School of Life Sciences, Fudan University, Shanghai, China, Ministry of Education Key Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan University, Shanghai, China, Center for genetic Epidemiology, School of Life Sciences, Fudan University, Shanghai, China, Center for Genetic Translational Medicine and Prevention, School of Public Health, Fudan University, Shanghai, China

Suqin Shen

Affiliation State Key Laboratory of Genetic Engineering, Collaborative Innovation Center for Genetics and Development, School of Life Sciences, Fudan University, Shanghai, China

He-Xi Ge Sai-Yin

Affiliation State Key Laboratory of Genetic Engineering, Collaborative Innovation Center for Genetics and Development, School of Life Sciences, Fudan University, Shanghai, China

Tao-Yang Chen

Affiliation Qidong Liver Cancer Institute, Qidong People’s Hospital, Qidong, Jiangsu, China

Pei-Xin Lu

Affiliation Qidong Liver Cancer Institute, Qidong People’s Hospital, Qidong, Jiangsu, China

Neng-Jin Wang

Affiliation Qidong Liver Cancer Institute, Qidong People’s Hospital, Qidong, Jiangsu, China

Weihua Ren

Affiliation Central Laboratory, Luoyang Central Hospital Affiliated to Zhengzhou University, Luoyang, Henan, China

Peng Huang

Affiliation College of Basic Medicine, Shanghai University of Medicine and Health Sciences, Shanghai, China

Jun Xie

Affiliation State Key Laboratory of Genetic Engineering, Collaborative Innovation Center for Genetics and Development, School of Life Sciences, Fudan University, Shanghai, China

Carly Conran

Affiliation Program for Personalized Cancer Care, NorthShore University HealthSystem, Pritzker School of Medicine, University of Chicago, Evanston, Illinois, United States of America

S. Lilly Zheng

Affiliation Program for Personalized Cancer Care, NorthShore University HealthSystem, Pritzker School of Medicine, University of Chicago, Evanston, Illinois, United States of America

Long Yu

Affiliation State Key Laboratory of Genetic Engineering, Collaborative Innovation Center for Genetics and Development, School of Life Sciences, Fudan University, Shanghai, China

Jianfeng Xu

Affiliations State Key Laboratory of Genetic Engineering, Collaborative Innovation Center for Genetics and Development, School of Life Sciences, Fudan University, Shanghai, China, Ministry of Education Key Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan University, Shanghai, China, Center for genetic Epidemiology, School of Life Sciences, Fudan University, Shanghai, China, Center for Genetic Translational Medicine and Prevention, School of Public Health, Fudan University, Shanghai, China, College of Basic Medicine, Shanghai University of Medicine and Health Sciences, Shanghai, China, Fudan Institute of Urology, Huashan Hospital, Fudan University, Shanghai, China

De-Ke Jiang

dekejiang@gmail.com

Affiliations State Key Laboratory of Genetic Engineering, Collaborative Innovation Center for Genetics and Development, School of Life Sciences, Fudan University, Shanghai, China, Ministry of Education Key Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan University, Shanghai, China, Center for genetic Epidemiology, School of Life Sciences, Fudan University, Shanghai, China, State Key Laboratory of Organ Failure Research, Guangdong Key Laboratory of Viral Hepatitis Research, Department of Infectious Diseases and Hepatology Unit, Nanfang Hospital, Southern Medical University, GuangZhou, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: DKJ LY JFX. Data curation: DKJ. Formal analysis: SZ GPY HTC. Funding acquisition: DKJ LY JFX. Investigation: DKJ SZ. Methodology: QYX ZQS XPM PYZ. Project administration: DKJ. Resources: TYC PXL NJW WHR HXGSY SQS PH JX. Writing – original draft: SZ. Writing – review & editing: DKJ CC SLZ.