About the Authors

Satsuki Tsuji

satsuki.may425@gmail.com

Affiliation Graduate School of Science and Technology, Ryukoku University, 1–5 Yokotani, Seta Oe-cho, Otsu, Japan

Masayuki Ushio

Affiliations Joint Research Center for Science and Technology, Ryukoku University, Otsu, Japan, Center for Ecological Research, Kyoto University, Hirano, Otsu, Japan, PRESTO, Japan Science and Technology Agency, Kawaguchi, Japan

Sho Sakurai

Affiliation Graduate School of Science and Technology, Ryukoku University, 1–5 Yokotani, Seta Oe-cho, Otsu, Japan

Toshifumi Minamoto

Affiliation Graduate School of Human Development and Environment, Kobe University, Tsurukabuto, Nada-ku, Kobe, Japan

Hiroki Yamanaka

Affiliation Faculty of Science and Technology, Ryukoku University, Yokotani, Seta Oe-cho, Otsu, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: ST TM HY. Formal analysis: ST MU SS. Investigation: ST. Writing – original draft: ST MU TM HY.